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Pristimantis  Jiménez de la Espada, 1870
Taxonomic Serial No.: 773300

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 Taxonomy and Nomenclature
       
  Kingdom: Animalia  
  Taxonomic Rank: Genus  
  Synonym(s): Trachyphrynus Goin and Cochran, 1963
  
    Cyclocephalus Jiménez de la Espada, 1875
  
    Hypodictyon Cope, 1885
  
    Pseudohyla Andersson, 1945
  
    Mucubatrachus La Marca, 2007
  
    Paramophrynella La Marca, 2007
  
  Common Name(s):    
       
  Taxonomic Status:    
  Current Standing: valid  
       
  Data Quality Indicators:    
  Record Credibility Rating: verified - standards met  
  Global Species Completeness: complete   
  Latest Record Review: 2019   
       

 Taxonomic Hierarchy
       
 KingdomAnimalia  – Animal, animaux, animals  
    SubkingdomBilateria  – triploblasts  
       InfrakingdomDeuterostomia   
          PhylumChordata  – cordés, cordado, chordates  
             SubphylumVertebrata  – vertebrado, vertébrés, vertebrates  
                InfraphylumGnathostomata   
                   SuperclassTetrapoda   
                      ClassAmphibia  – amphibiens, anfíbio, Amphibians  
                         OrderAnura  – Frogs, perereca, rã, sapo, crapauds, grenouilles, Toads  
                            SuperfamilyBrachycephaloidea Günther, 1858  
                               FamilyCraugastoridae Hedges, Duellman and Heinicke, 2008  
                                  SubfamilyCeuthomantinae Heinicke, Duellman, Trueb, Means, MacCulloch and Hedges, 2009  
                                     GenusPristimantis Jiménez de la Espada, 1870  
    Direct Children:  
                                        Species Pristimantis aaptus (Lynch and Lescure, 1980)  
                                        Species Pristimantis abakapa Rojas-Runjaic, Salerno, Señaris and Pauly, 2013  
                                        Species Pristimantis academicus Lehr, Moravec and Gagliardi-Urrutia, 2010  
                                        Species Pristimantis acatallelus (Lynch and Ruiz-Carranza, 1983)  
                                        Species Pristimantis acerus (Lynch and Duellman, 1980)  
                                        Species Pristimantis achatinus (Boulenger, 1898)  
                                        Species Pristimantis actinolaimus (Lynch and Rueda-Almonacid, 1998)  
                                        Species Pristimantis actites (Lynch, 1979)  
                                        Species Pristimantis acuminatus (Shreve, 1935)  
                                        Species Pristimantis acutirostris (Lynch, 1984)  
                                        Species Pristimantis adiastolus Duellman and Hedges, 2007  
                                        Species Pristimantis adnus Crawford, Ryan and Jaramillo, 2010  
                                        Species Pristimantis aemulatus (Ruiz-Carranza, Lynch and Ardila-Robayo, 1997)  
                                        Species Pristimantis affinis (Werner, 1899)  
                                        Species Pristimantis alalocophus (Roa-Trujillo and Ruiz-Carranza, 1991)  
                                        Species Pristimantis albericoi (Lynch and Ruiz-Carranza, 1996)  
                                        Species Pristimantis albertus Duellman and Hedges, 2007  
                                        Species Pristimantis albujai Brito-M., Batallas-R. and Yánez-Muñoz, 2017  
                                        Species Pristimantis allpapuyu Yánez-Muñoz, Sánchez-Nivicela and Reyes-Puig, 2016  
                                        Species Pristimantis almendariz Brito-M. and Pozo-Zamora, 2013  
                                        Species Pristimantis altae (Dunn, 1942)  
                                        Species Pristimantis altamazonicus (Barbour and Dunn, 1921)  
                                        Species Pristimantis altamnis Elmer and Cannatella, 2008  
                                        Species Pristimantis ameliae Barrio-Amorós, 2012  
                                        Species Pristimantis amydrotus (Duellman and Lehr, 2007)  
                                        Species Pristimantis andinogigas Yánez-Muñoz, Veintimilla-Yánez, Batallas-R. and Cisneros-Heredia, 2019  
                                        Species Pristimantis andinognomus Lehr and Coloma, 2008  
                                        Species Pristimantis anemerus (Duellman and Pramuk, 1999)  
                                        Species Pristimantis angustilineatus (Lynch, 1998)  
                                        Species Pristimantis aniptopalmatus (Duellman and Hedges, 2005)  
                                        Species Pristimantis anolirex (Lynch, 1983)  
                                        Species Pristimantis anotis (Walker and Test, 1955)  
                                        Species Pristimantis antisuyu Catenazzi and Lehr, 2018  
                                        Species Pristimantis apiculatus (Lynch and Burrowes, 1990)  
                                        Species Pristimantis appendiculatus (Werner, 1894)  
                                        Species Pristimantis aquilonaris Lehr, Aguilar, Siu-Ting and Jordán, 2007  
                                        Species Pristimantis ardalonychus (Duellman and Pramuk, 1999)  
                                        Species Pristimantis ardyae Reyes-Puig, Reyes-Puig and Yánez-Muñoz, 2013  
                                        Species Pristimantis ashaninka Lehr and Moravec, 2017  
                                        Species Pristimantis atrabracus (Duellman and Pramuk, 1999)  
                                        Species Pristimantis atratus (Lynch, 1979)  
                                        Species Pristimantis attenboroughi Lehr and von May, 2017  
                                        Species Pristimantis aurantiguttatus (Ruiz-Carranza, Lynch and Ardila-Robayo, 1997)  
                                        Species Pristimantis aureolineatus (Guayasamin, Ron, Cisneros-Heredia, Lamar and McCracken, 2006)  
                                        Species Pristimantis aureoventris Kok, Means and Bossuyt, 2011  
                                        Species Pristimantis auricarens (Myers and Donnelly, 2008)  
                                        Species Pristimantis avicuporum (Duellman and Pramuk, 1999)  
                                        Species Pristimantis avius (Myers and Donnelly, 1997)  
                                        Species Pristimantis bacchus (Lynch, 1984)  
                                        Species Pristimantis baiotis (Lynch, 1998)  
                                        Species Pristimantis balionotus (Lynch, 1979)  
                                        Species Pristimantis bambu Arteaga-Navarro and Guayasamin, 2011  
                                        Species Pristimantis barrigai Brito and Almendáriz C., 2018  
                                        Species Pristimantis baryecuus (Lynch, 1979)  
                                        Species Pristimantis batrachites (Lynch, 2003)  
                                        Species Pristimantis bearsei (Duellman, 1992)  
                                        Species Pristimantis bellae Reyes-Puig and Yánez-Muñoz, 2012  
                                        Species Pristimantis bellator Lehr, Aguilar, Siu-Ting and Jordán, 2007  
                                        Species Pristimantis bellona (Lynch, 1992)  
                                        Species Pristimantis bernali (Lynch, 1986)  
                                        Species Pristimantis bicantus Guayasamin and Funk, 2009  
                                        Species Pristimantis bicolor (Rueda-Almonacid and Lynch, 1983)  
                                        Species Pristimantis bicumulus (Peters, 1863)  
                                        Species Pristimantis bipunctatus (Duellman and Hedges, 2005)  
                                        Species Pristimantis boconoensis (Rivero and Mayorga, 1973)  
                                        Species Pristimantis bogotensis (Peters, 1863)  
                                        Species Pristimantis boucephalus Lehr, Moravec, Cusi and Gvozdík, 2017  
                                        Species Pristimantis boulengeri (Lynch, 1981)  
                                        Species Pristimantis bounides Lehr, von May, Moravec and Cusi, 2017  
                                        Species Pristimantis brevicrus (Andersson, 1945)  
                                        Species Pristimantis brevifrons (Lynch, 1981)  
                                        Species Pristimantis briceni (Boulenger, 1903)  
                                        Species Pristimantis bromeliaceus (Lynch, 1979)  
                                        Species Pristimantis buccinator (Rodríguez, 1994)  
                                        Species Pristimantis buckleyi (Boulenger, 1882)  
                                        Species Pristimantis buenaventura Arteaga-Navarro, Pyron, Peñafiel, Romero-Barreto, Culebras, Bustamante, Yánez-Muñoz et al., 2016  
                                        Species Pristimantis bustamante Chaparro, Motta, Gutiérrez and Padial, 2012  
                                        Species Pristimantis cabrerai (Cochran and Goin, 1970)  
                                        Species Pristimantis cacao (Lynch, 1992)  
                                        Species Pristimantis caeruleonotus Lehr, Aguilar, Siu-Ting and Jordán, 2007  
                                        Species Pristimantis cajamarcensis (Barbour and Noble, 1920)  
                                        Species Pristimantis calcaratus (Boulenger, 1908)  
                                        Species Pristimantis calcarulatus (Lynch, 1976)  
                                        Species Pristimantis calima Ospina-Sarria and Duellman, 2019  
                                        Species Pristimantis caniari Rámírez-Jaramillo, Reyes-Puig, Batallas-R. and Yánez-Muñoz, 2018  
                                        Species Pristimantis cantitans (Myers and Donnelly, 1996)  
                                        Species Pristimantis capitonis (Lynch, 1998)  
                                        Species Pristimantis caprifer (Lynch, 1977)  
                                        Species Pristimantis carlosceroni Valencia, Bejarano-Muñoz and Yánez-Muñoz, 2013  
                                        Species Pristimantis carlossanchezi (Arroyo, 2007)  
                                        Species Pristimantis carmelitae (Ruthven, 1922)  
                                        Species Pristimantis carranguerorum (Lynch, 1994)  
                                        Species Pristimantis carvalhoi (Lutz in Lutz and Kloss, 1952)  
                                        Species Pristimantis caryophyllaceus (Barbour, 1928)  
                                        Species Pristimantis cedros Hutter and Guayasamin, 2015  
                                        Species Pristimantis celator (Lynch, 1976)  
                                        Species Pristimantis cerasinus (Cope, 1875)  
                                        Species Pristimantis ceuthospilus (Duellman and Wild, 1993)  
                                        Species Pristimantis chalceus (Peters, 1873)  
                                        Species Pristimantis charlottevillensis (Kaiser, Dwyer, Feichtinger and Schmid, 1995)  
                                        Species Pristimantis chiastonotus (Lynch and Hoogmoed, 1977)  
                                        Species Pristimantis chimu Lehr, 2007  
                                        Species Pristimantis chloronotus (Lynch, 1969)  
                                        Species Pristimantis chrysops (Lynch and Ruiz-Carranza, 1996)  
                                        Species Pristimantis churuwiai Brito-M., Batallas-R. and Yánez-Muñoz, 2017  
                                        Species Pristimantis citriogaster (Duellman, 1992)  
                                        Species Pristimantis colodactylus (Lynch, 1979)  
                                        Species Pristimantis colomai (Lynch and Duellman, 1997)  
                                        Species Pristimantis colonensis (Mueses-Cisneros, 2007)  
                                        Species Pristimantis colostichos (La Marca and Smith, 1982)  
                                        Species Pristimantis condor (Lynch and Duellman, 1980)  
                                        Species Pristimantis conservatio Barrio-Amorós, Heinicke and Hedges, 2013  
                                        Species Pristimantis conspicillatus (Günther, 1858)  
                                        Species Pristimantis cordovae (Lehr and Duellman, 2007)  
                                        Species Pristimantis corniger (Lynch and Suárez-Mayorga, 2003)  
                                        Species Pristimantis coronatus Lehr and Duellman, 2007  
                                        Species Pristimantis corrugatus (Duellman, Lehr and Venegas, 2006)  
                                        Species Pristimantis cosnipatae (Duellman, 1978)  
                                        Species Pristimantis cremnobates (Lynch and Duellman, 1980)  
                                        Species Pristimantis crenunguis (Lynch, 1976)  
                                        Species Pristimantis cristinae (Lynch and Ruiz-Carranza, 1985)  
                                        Species Pristimantis croceoinguinis (Lynch, 1968)  
                                        Species Pristimantis crucifer (Boulenger, 1899)  
                                        Species Pristimantis cruciocularis (Lehr, Lundberg, Aguilar and von May, 2006)  
                                        Species Pristimantis cruentus (Peters, 1873)  
                                        Species Pristimantis cryophilius (Lynch, 1979)  
                                        Species Pristimantis cryptomelas (Lynch, 1979)  
                                        Species Pristimantis cuentasi (Lynch, 2003)  
                                        Species Pristimantis culatensis (La Marca, 2007)  
                                        Species Pristimantis cuneirostris (Duellman and Pramuk, 1999)  
                                        Species Pristimantis curtipes (Boulenger, 1882)  
                                        Species Pristimantis danae (Duellman, 1978)  
                                        Species Pristimantis degener (Lynch and Duellman, 1997)  
                                        Species Pristimantis deinops (Lynch, 1996)  
                                        Species Pristimantis delicatus (Ruthven, 1917)  
                                        Species Pristimantis delius (Duellman and Mendelson, 1995)  
                                        Species Pristimantis dendrobatoides Means and Savage, 2007  
                                        Species Pristimantis devillei (Boulenger, 1880)  
                                        Species Pristimantis deyi Lehr, Gregory and Catenazzi, 2013  
                                        Species Pristimantis diadematus (Jiménez de la Espada, 1875)  
                                        Species Pristimantis diaphonus (Lynch, 1986)  
                                        Species Pristimantis diogenes (Lynch and Ruiz-Carranza, 1996)  
                                        Species Pristimantis dissimulatus (Lynch and Duellman, 1997)  
                                        Species Pristimantis divnae Lehr and von May, 2009  
                                        Species Pristimantis dorado Rivera-Correa, Lamadrid-Feris and Crawford, 2016  
                                        Species Pristimantis dorsopictus (Rivero and Serna, 1988)  
                                        Species Pristimantis duellmani (Lynch, 1980)  
                                        Species Pristimantis duende (Lynch, 2001)  
                                        Species Pristimantis dundeei (Heyer and Muñoz, 1999)  
                                        Species Pristimantis ecuadorensis Guayasamin, Hutter, Tapia, Culebras, Peñafiel, Pyron, Morochz, Funk and Arteaga-Navarro, 2017  
                                        Species Pristimantis educatoris Ryan, Lips and Giermakowski, 2010  
                                        Species Pristimantis elegans (Peters, 1863)  
                                        Species Pristimantis enigmaticus Ortega-Andrade, Rojas-Soto, Valencia, Espinosa de los Monteros, Morrone, Ron and Cannatella, 2015  
                                        Species Pristimantis epacrus (Lynch and Suárez-Mayorga, 2000)  
                                        Species Pristimantis eremitus (Lynch, 1980)  
                                        Species Pristimantis eriphus (Lynch and Duellman, 1980)  
                                        Species Pristimantis ernesti (Flores, 1987)  
                                        Species Pristimantis erythroinguinis Catenazzi and Lehr, 2018  
                                        Species Pristimantis erythropleura (Boulenger, 1896)  
                                        Species Pristimantis erythros Sánchez-Nivicela, Celi-Piedra, Posse-Sarmiento, Urgilés, Yánez-Muñoz and Cisneros-Heredia, 2018  
                                        Species Pristimantis esmeraldas (Guayasamin, 2004)  
                                        Species Pristimantis espedeus Fouquet, Martinez, Courtois, Dewynter, Pineau, Gaucher, Blanc, Marty and Kok, 2013  
                                        Species Pristimantis eugeniae (Lynch and Duellman, 1997)  
                                        Species Pristimantis euphronides (Schwartz, 1967)  
                                        Species Pristimantis eurydactylus (Hedges and Schlüter, 1992)  
                                        Species Pristimantis exoristus (Duellman and Pramuk, 1999)  
                                        Species Pristimantis factiosus (Lynch and Rueda-Almonacid, 1998)  
                                        Species Pristimantis fallax (Lynch and Rueda-Almonacid, 1999)  
                                        Species Pristimantis farisorum Mueses-Cisneros, Perdomo-Castillo and Cepeda-Quilindo, 2013  
                                        Species Pristimantis fasciatus Barrio-Amorós, Rojas-Runjaic and Infante-Rivero, 2008  
                                        Species Pristimantis fenestratus (Steindachner, 1864)  
                                        Species Pristimantis festae (Peracca, 1904)  
                                        Species Pristimantis fetosus (Lynch and Rueda-Almonacid, 1998)  
                                        Species Pristimantis flabellidiscus (La Marca, 2007)  
                                        Species Pristimantis floridus (Lynch and Duellman, 1997)  
                                        Species Pristimantis frater (Werner, 1899)  
                                        Species Pristimantis gagliardoi Bustamante and Mendelson, 2008  
                                        Species Pristimantis gaigei (Dunn, 1931)  
                                        Species Pristimantis galdi Jiménez de la Espada, 1870  
                                        Species Pristimantis ganonotus (Duellman and Lynch, 1988)  
                                        Species Pristimantis geminus Kaiser, Barrio-Amorós, Rivas-Fuenmayor, Steinlein and Schmid, 2015  
                                        Species Pristimantis gentryi (Lynch and Duellman, 1997)  
                                        Species Pristimantis ginesi (Rivero, 1964)  
                                        Species Pristimantis gladiator (Lynch, 1976)  
                                        Species Pristimantis glandulosus (Boulenger, 1880)  
                                        Species Pristimantis gracilis (Lynch, 1986)  
                                        Species Pristimantis gralarias Guayasamin, Arteaga-Navarro and Hutter, 2018  
                                        Species Pristimantis grandiceps (Lynch, 1984)  
                                        Species Pristimantis gryllus Barrio-Amorós, Guayasamin and Hedges, 2012  
                                        Species Pristimantis guaiquinimensis (Schlüter and Rödder, 2007)  
                                        Species Pristimantis gualacenio Urgilés, Sánchez-Nivicela, Nieves and Yánez-Muñoz, 2014  
                                        Species Pristimantis gutturalis (Hoogmoed, Lynch and Lescure, 1977)  
                                        Species Pristimantis hamiotae (Flores, 1994)  
                                        Species Pristimantis hampatusami Yánez-Muñoz, Sánchez-Nivicela and Reyes-Puig, 2016  
                                        Species Pristimantis hectus (Lynch and Burrowes, 1990)  
                                        Species Pristimantis helvolus (Lynch and Rueda-Almonacid, 1998)  
                                        Species Pristimantis hernandezi (Lynch and Ruiz-Carranza, 1983)  
                                        Species Pristimantis hoogmoedi Kaiser, Barrio-Amorós, Rivas-Fuenmayor, Steinlein and Schmid, 2015  
                                        Species Pristimantis huicundo (Guayasamin, Almeida-Reinoso and Nogales-Sornosa, 2004)  
                                        Species Pristimantis humboldti Lehr, von May, Moravec and Cusi, 2017  
                                        Species Pristimantis hybotragus (Lynch, 1992)  
                                        Species Pristimantis ignicolor (Lynch and Duellman, 1980)  
                                        Species Pristimantis iiap Padial, Gagliardi-Urrutia, Chaparro and Gutiérrez, 2016  
                                        Species Pristimantis illotus (Lynch and Duellman, 1997)  
                                        Species Pristimantis imitatrix (Duellman, 1978)  
                                        Species Pristimantis imthurni Kok, 2013  
                                        Species Pristimantis incanus (Lynch and Duellman, 1980)  
                                        Species Pristimantis incertus (Lutz, 1927)  
                                        Species Pristimantis incomptus (Lynch and Duellman, 1980)  
                                        Species Pristimantis infraguttatus (Duellman and Pramuk, 1999)  
                                        Species Pristimantis inguinalis (Parker, 1940)  
                                        Species Pristimantis insignitus (Ruthven, 1917)  
                                        Species Pristimantis inusitatus (Lynch and Duellman, 1980)  
                                        Species Pristimantis ixalus (Lynch, 2003)  
                                        Species Pristimantis jabonensis (La Marca, 2007)  
                                        Species Pristimantis jaguensis Rivera-Prieto, Rivera-Correa and Daza-R., 2014  
                                        Species Pristimantis jaimei (Lynch, 1992)  
                                        Species Pristimantis jamescameroni Kok, 2013  
                                        Species Pristimantis jester Means and Savage, 2007  
                                        Species Pristimantis johannesdei (Rivero and Serna, 1988)  
                                        Species Pristimantis jorgevelosai (Lynch, 1994)  
                                        Species Pristimantis juanchoi (Lynch, 1996)  
                                        Species Pristimantis jubatus (García and Lynch, 2006)  
                                        Species Pristimantis kareliae (La Marca, 2005)  
                                        Species Pristimantis katoptroides (Flores, 1988)  
                                        Species Pristimantis kelephus (Lynch, 1998)  
                                        Species Pristimantis kichwarum Elmer and Cannatella, 2008  
                                        Species Pristimantis kirklandi (Flores, 1985)  
                                        Species Pristimantis koehleri Padial and De la Riva, 2009  
                                        Species Pristimantis kuri Yánez-Muñoz, Sánchez-Nivicela and Reyes-Puig, 2016  
                                        Species Pristimantis labiosus (Lynch, Ruiz-Carranza and Ardila-Robayo, 1994)  
                                        Species Pristimantis lacrimosus (Jiménez de la Espada, 1875)  
                                        Species Pristimantis lancinii (Donoso-Barros, 1965)  
                                        Species Pristimantis lanthanites (Lynch, 1975)  
                                        Species Pristimantis lasalleorum (Lynch, 1995)  
                                        Species Pristimantis lassoalcalai Barrio-Amorós, Rojas-Runjaic and Barros, 2010  
                                        Species Pristimantis latericius Batallas-R. and Brito-M., 2014  
                                        Species Pristimantis laticlavius (Lynch and Burrowes, 1990)  
                                        Species Pristimantis latidiscus (Boulenger, 1898)  
                                        Species Pristimantis latro Oliveira, Rodrigues, Kaefer, Pinto and Hernández-Ruz, 2017  
                                        Species Pristimantis lemur (Lynch and Rueda-Almonacid, 1998)  
                                        Species Pristimantis leoni (Lynch, 1976)  
                                        Species Pristimantis leopardus Rivera-Correa, Jiménez-Rivillas and Daza-R., 2017  
                                        Species Pristimantis leptolophus (Lynch, 1980)  
                                        Species Pristimantis leucopus (Lynch, 1976)  
                                        Species Pristimantis leucorrhinus Boano, Mazzotti and Sindaco, 2008  
                                        Species Pristimantis librarius (Flores and Vigle, 1994)  
                                        Species Pristimantis lichenoides (Lynch and Rueda-Almonacid, 1997)  
                                        Species Pristimantis limoncochensis Ortega-Andrade, Rojas-Soto, Valencia, Espinosa de los Monteros, Morrone, Ron and Cannatella, 2015  
                                        Species Pristimantis lindae (Duellman, 1978)  
                                        Species Pristimantis lirellus (Dwyer, 1995)  
                                        Species Pristimantis lividus (Lynch and Duellman, 1980)  
                                        Species Pristimantis llanganati Navarrete, Venegas and Ron, 2016  
                                        Species Pristimantis llojsintuta (Köhler and Lötters, 1999)  
                                        Species Pristimantis longicorpus Kaiser, Barrio-Amorós, Rivas-Fuenmayor, Steinlein and Schmid, 2015  
                                        Species Pristimantis loujosti Yánez-Muñoz, Cisneros-Heredia and Reyes-Puig, 2011  
                                        Species Pristimantis loustes (Lynch, 1979)  
                                        Species Pristimantis lucasi Duellman and Chaparro, 2008  
                                        Species Pristimantis lucidosignatus Rödder and Schmitz, 2009  
                                        Species Pristimantis luscombei (Duellman and Mendelson, 1995)  
                                        Species Pristimantis luteolateralis (Lynch, 1976)  
                                        Species Pristimantis lutitus (Lynch, 1984)  
                                        Species Pristimantis lymani (Barbour and Noble, 1920)  
                                        Species Pristimantis lynchi (Duellman and Simmons, 1977)  
                                        Species Pristimantis lythrodes (Lynch and Lescure, 1980)  
                                        Species Pristimantis macrummendozai Acosta-Galvis, 2015  
                                        Species Pristimantis maculosus (Lynch, 1991)  
                                        Species Pristimantis malkini (Lynch, 1980)  
                                        Species Pristimantis mallii Reyes-Puig, Reyes-Puig, Velarde-Garcéz, Dávalos, Mancero, Navarrete, Yánez-Muñoz et al., 2019  
                                        Species Pristimantis marahuaka (Fuentes-Ramos and Barrio-Amorós, 2004)  
                                        Species Pristimantis marcoreyesi Reyes-Puig, Reyes-Puig, Rámirez-Jaramillo, Pérez-L. and Yánez-Munoz, 2015  
                                        Species Pristimantis mariaelenae Venegas and Duellman, 2012  
                                        Species Pristimantis marmoratus (Boulenger, 1900)  
                                        Species Pristimantis mars (Lynch and Ruiz-Carranza, 1996)  
                                        Species Pristimantis martiae (Lynch, 1974)  
                                        Species Pristimantis matidiktyo Ortega-Andrade and Valencia, 2012  
                                        Species Pristimantis mazar Guayasamin and Arteaga-Navarro, 2013  
                                        Species Pristimantis medemi (Lynch, 1994)  
                                        Species Pristimantis megalops (Ruthven, 1917)  
                                        Species Pristimantis melanogaster (Duellman and Pramuk, 1999)  
                                        Species Pristimantis melanoproctus (Rivero, 1984)  
                                        Species Pristimantis memorans (Myers and Donnelly, 1997)  
                                        Species Pristimantis mendax (Duellman, 1978)  
                                        Species Pristimantis meridionalis (Lehr and Duellman, 2007)  
                                        Species Pristimantis merostictus (Lynch, 1984)  
                                        Species Pristimantis metabates (Duellman and Pramuk, 1999)  
                                        Species Pristimantis miktos Ortego-Andrade and Venegas, 2014  
                                        Species Pristimantis mindo Arteaga-Navarro, Yáñez-Muñoz and Guayasamin, 2013  
                                        Species Pristimantis minimus Terán-Valdez and Guayasamin, 2010  
                                        Species Pristimantis minutulus Duellman and Hedges, 2007  
                                        Species Pristimantis miyatai (Lynch, 1984)  
                                        Species Pristimantis mnionaetes (Lynch, 1998)  
                                        Species Pristimantis modipeplus (Lynch, 1981)  
                                        Species Pristimantis molybrignus (Lynch, 1986)  
                                        Species Pristimantis mondolfii (Rivero, 1984)  
                                        Species Pristimantis moro (Savage, 1965)  
                                        Species Pristimantis muchimuk Barrio-Amorós, Mesa, Brewer-Carías and McDiarmid, 2010  
                                        Species Pristimantis munozi Rojas-Runjaic, Delgado C. and Guayasamin, 2014  
                                        Species Pristimantis muranunka Brito M., Almendariz-C., Batallas R. and Ron, 2017  
                                        Species Pristimantis muricatus (Lynch and Miyata, 1980)  
                                        Species Pristimantis muscosus (Duellman and Pramuk, 1999)  
                                        Species Pristimantis museosus (Ibáñez, Jaramillo and Arosemena, 1994)  
                                        Species Pristimantis mutabilis Guayasamin, Krynak, Krynak, Culebras and Hutter, 2015  
                                        Species Pristimantis myersi (Goin and Cochran, 1963)  
                                        Species Pristimantis myops (Lynch, 1998)  
                                        Species Pristimantis nebulosus (Henle, 1992)  
                                        Species Pristimantis nephophilus (Duellman and Pramuk, 1999)  
                                        Species Pristimantis nervicus (Lynch, 1994)  
                                        Species Pristimantis nicefori (Cochran and Goin, 1970)  
                                        Species Pristimantis nietoi Arteaga-Navarro, Pyron, Peñafiel, Romero-Barreto, Culebras, Bustamante, Yánez-Muñoz et al., 2016  
                                        Species Pristimantis nigrogriseus (Andersson, 1945)  
                                        Species Pristimantis nimbus Urgilés, Posse, Timbe, Astudillo and Sánchez-Nivicela, 2017  
                                        Species Pristimantis nubisilva Kaiser, Barrio-Amorós, Rivas-Fuenmayor, Steinlein and Schmid, 2015  
                                        Species Pristimantis nyctophylax (Lynch, 1976)  
                                        Species Pristimantis obmutescens (Lynch, 1980)  
                                        Species Pristimantis ocellatus (Lynch and Burrowes, 1990)  
                                        Species Pristimantis ockendeni (Boulenger, 1912)  
                                        Species Pristimantis ocreatus (Lynch, 1981)  
                                        Species Pristimantis olivaceus (Köhler, Morales, Lötters, Reichle and Aparicio, 1998)  
                                        Species Pristimantis omeviridis Ortega-Andrade, Rojas-Soto, Valencia, Espinosa de los Monteros, Morrone, Ron and Cannatella, 2015  
                                        Species Pristimantis onorei Rödder and Schmitz, 2009  
                                        Species Pristimantis orcesi (Lynch, 1972)  
                                        Species Pristimantis orcus Lehr, Catenazzi and Rodríguez, 2009  
                                        Species Pristimantis orestes (Lynch, 1979)  
                                        Species Pristimantis ornatissimus (Despax, 1911)  
                                        Species Pristimantis ornatus (Lehr, Lundberg, Aguilar and von May, 2006)  
                                        Species Pristimantis orpacobates (Lynch, Ruiz-Carranza and Ardila-Robayo, 1994)  
                                        Species Pristimantis orphnolaimus (Lynch, 1970)  
                                        Species Pristimantis ortizi (Guayasamin, Almeida-Reinoso and Nogales-Sornosa, 2004)  
                                        Species Pristimantis padiali Moravec, Lehr, Perez-Peña, Lopez, Gagliardi-Urrutia and Arista-Tuanama, 2010  
                                        Species Pristimantis padrecarlosi (Mueses-Cisneros, 2006)  
                                        Species Pristimantis pahuma Hutter and Guayasamin, 2015  
                                        Species Pristimantis paisa (Lynch and Ardila-Robayo, 1999)  
                                        Species Pristimantis palmeri (Boulenger, 1912)  
                                        Species Pristimantis paquishae Brito-M., Batallas-R. and Velalcázar, 2014  
                                        Species Pristimantis paramerus (Rivero, 1984)  
                                        Species Pristimantis pardalinus (Lehr, Lundberg, Aguilar and von May, 2006)  
                                        Species Pristimantis pardalis (Barbour, 1928)  
                                        Species Pristimantis parectatus (Lynch and Rueda-Almonacid, 1998)  
                                        Species Pristimantis pariagnomus Kaiser, Barrio-Amorós, Rivas-Fuenmayor, Steinlein and Schmid, 2015  
                                        Species Pristimantis parvillus (Lynch, 1976)  
                                        Species Pristimantis pastazensis (Andersson, 1945)  
                                        Species Pristimantis pataikos (Duellman and Pramuk, 1999)  
                                        Species Pristimantis paulodutrai (Bokermann, 1975)  
                                        Species Pristimantis paululus (Lynch, 1974)  
                                        Species Pristimantis pecki (Duellman and Lynch, 1988)  
                                        Species Pristimantis pedimontanus (La Marca, 2004)  
                                        Species Pristimantis penelopus (Lynch and Rueda-Almonacid, 1999)  
                                        Species Pristimantis peraticus (Lynch, 1980)  
                                        Species Pristimantis percnopterus (Duellman and Pramuk, 1999)  
                                        Species Pristimantis percultus (Lynch, 1979)  
                                        Species Pristimantis permixtus (Lynch, Ruiz-Carranza and Ardila-Robayo, 1994)  
                                        Species Pristimantis peruvianus (Melin, 1941)  
                                        Species Pristimantis petersi (Lynch and Duellman, 1980)  
                                        Species Pristimantis petrobardus (Duellman, 1991)  
                                        Species Pristimantis phalaroinguinis (Duellman and Lehr, 2007)  
                                        Species Pristimantis phalarus (Lynch, 1998)  
                                        Species Pristimantis pharangobates (Duellman, 1978)  
                                        Species Pristimantis philipi (Lynch and Duellman, 1995)  
                                        Species Pristimantis phoxocephalus (Lynch, 1979)  
                                        Species Pristimantis phragmipleuron (Rivero and Serna, 1988)  
                                        Species Pristimantis piceus (Lynch, Ruiz-Carranza and Ardila-Robayo, 1996)  
                                        Species Pristimantis pichincha Yánez-Muñoz, Reyes-Puig, Bejarano-Muñoz and Ron, 2016  
                                        Species Pristimantis pinchaque Reyes-Puig, Reyes-Puig, Pérez-L. and Yánez-Muñoz, 2015  
                                        Species Pristimantis pinguis (Duellman and Pramuk, 1999)  
                                        Species Pristimantis pirrensis (Ibáñez and Crawford, 2004)  
                                        Species Pristimantis platychilus (Lynch, 1996)  
                                        Species Pristimantis platydactylus (Boulenger, 1903)  
                                        Species Pristimantis pleurostriatus (Rivero, 1984)  
                                        Species Pristimantis pluvialis Shepack, von May, Ttito and Catenazzi, 2016  
                                        Species Pristimantis polemistes (Lynch and Ardila-Robayo, 2004)  
                                        Species Pristimantis polychrus (Ruiz-Carranza, Lynch and Ardila-Robayo, 1997)  
                                        Species Pristimantis prolatus (Lynch and Duellman, 1980)  
                                        Species Pristimantis prometeii Székely, Cogalniceanu, Székely, Páez and Ron, 2016  
                                        Species Pristimantis proserpens (Lynch, 1979)  
                                        Species Pristimantis pruinatus (Myers and Donnelly, 1996)  
                                        Species Pristimantis pseudoacuminatus (Shreve, 1935)  
                                        Species Pristimantis pteridophilus (Lynch and Duellman, 1997)  
                                        Species Pristimantis ptochus (Lynch, 1998)  
                                        Species Pristimantis pugnax (Lynch, 1973)  
                                        Species Pristimantis puipui Lehr, von May, Moravec and Cusi, 2017  
                                        Species Pristimantis pulchridormientes Chávez and Catenazzi, 2016  
                                        Species Pristimantis pulvinatus (Rivero, 1968)  
                                        Species Pristimantis punzan Reyes-Puig, Reyes-Puig, Rámirez-Jaramillo, Pérez-L. and Yánez-Munoz, 2015  
                                        Species Pristimantis puruscafeum Reyes-Puig, Reyes-Puig, Rámirez-Jaramillo, Pérez-L. and Yánez-Munoz, 2015  
                                        Species Pristimantis pycnodermis (Lynch, 1979)  
                                        Species Pristimantis pyrrhomerus (Lynch, 1976)  
                                        Species Pristimantis quantus (Lynch, 1998)  
                                        Species Pristimantis quaquaversus (Lynch, 1974)  
                                        Species Pristimantis quicato Ospina-Sarria, Méndez-Narváez, Burbano-Yandi and Bolívar-García, 2011  
                                        Species Pristimantis quinquagesimus (Lynch and Trueb, 1980)  
                                        Species Pristimantis racemus (Lynch, 1980)  
                                        Species Pristimantis ramagii (Boulenger, 1888)  
                                        Species Pristimantis reclusus (Lynch, 2003)  
                                        Species Pristimantis reichlei Padial and De la Riva, 2009  
                                        Species Pristimantis renjiforum (Lynch, 2000)  
                                        Species Pristimantis repens (Lynch, 1984)  
                                        Species Pristimantis restrepoi (Lynch, 1996)  
                                        Species Pristimantis reticulatus (Walker and Test, 1955)  
                                        Species Pristimantis rhabdocnemus (Duellman and Hedges, 2005)  
                                        Species Pristimantis rhabdolaemus (Duellman, 1978)  
                                        Species Pristimantis rhigophilus (La Marca, 2007)  
                                        Species Pristimantis rhodoplichus (Duellman and Wild, 1993)  
                                        Species Pristimantis rhodostichus (Duellman and Pramuk, 1999)  
                                        Species Pristimantis ridens (Cope, 1866)  
                                        Species Pristimantis rivasi Barrio-Amorós, Rojas-Runjaic and Barros, 2010  
                                        Species Pristimantis riveroi (Lynch and La Marca, 1993)  
                                        Species Pristimantis riveti (Despax, 1911)  
                                        Species Pristimantis romanorum Yánez-Muñoz, Meza-Ramos, Cisneros-Heredia and Reyes-Puig, 2011  
                                        Species Pristimantis roni Yanez-Munoz, Bejarano-Munoz, Brito-M. and Batallas-R., 2014  
                                        Species Pristimantis rosadoi (Flores, 1988)  
                                        Species Pristimantis roseus (Boulenger, 1918)  
                                        Species Pristimantis royi (Morales, 2007)  
                                        Species Pristimantis rozei (Rivero, 1961)  
                                        Species Pristimantis rubicundus (Jiménez de la Espada, 1875)  
                                        Species Pristimantis ruedai (Ruiz-Carranza, Lynch and Ardila-Robayo, 1997)  
                                        Species Pristimantis rufioculis (Duellman and Pramuk, 1999)  
                                        Species Pristimantis rufoviridis Valencia, Yánez-Muñoz, Betancourt-Yépez, Terán-Valdez and Guayasamin, 2011  
                                        Species Pristimantis ruidus (Lynch, 1979)  
                                        Species Pristimantis ruthveni (Lynch and Ruiz-Carranza, 1985)  
                                        Species Pristimantis sacharuna Reyes-Puig, Reyes-Puig, Pérez-L. and Yánez-Muñoz, 2015  
                                        Species Pristimantis sagittulus (Lehr, Aguilar and Duellman, 2004)  
                                        Species Pristimantis salaputium (Duellman, 1978)  
                                        Species Pristimantis saltissimus Means and Savage, 2007  
                                        Species Pristimantis samaipatae (Köhler and Jungfer, 1995)  
                                        Species Pristimantis sambalan Brito-M., Batallas-R. and Yánez-Muñoz, 2017  
                                        Species Pristimantis sanctaemartae (Ruthven, 1917)  
                                        Species Pristimantis sanguineus (Lynch, 1998)  
                                        Species Pristimantis sarisarinama Barrio-Amorós and Brewer-Carias, 2008  
                                        Species Pristimantis satagius (Lynch, 1995)  
                                        Species Pristimantis saturninoi Brito-M., Batallas-R. and Yánez-Muñoz, 2017  
                                        Species Pristimantis savagei (Pyburn and Lynch, 1981)  
                                        Species Pristimantis schultei (Duellman, 1990)  
                                        Species Pristimantis scitulus (Duellman, 1978)  
                                        Species Pristimantis scoloblepharus (Lynch, 1991)  
                                        Species Pristimantis scolodiscus (Lynch and Burrowes, 1990)  
                                        Species Pristimantis scopaeus (Lynch, Ruiz-Carranza and Ardila-Robayo, 1996)  
                                        Species Pristimantis seorsus Lehr, 2007  
                                        Species Pristimantis serendipitus (Duellman and Pramuk, 1999)  
                                        Species Pristimantis shrevei (Schwartz, 1967)  
                                        Species Pristimantis signifer (Ruiz-Carranza, Lynch and Ardila-Robayo, 1997)  
                                        Species Pristimantis silverstonei (Lynch and Ruiz-Carranza, 1996)  
                                        Species Pristimantis simonbolivari (Wiens and Coloma, 1992)  
                                        Species Pristimantis simonsii (Boulenger, 1900)  
                                        Species Pristimantis simoteriscus (Lynch, Ruiz-Carranza and Ardila-Robayo, 1997)  
                                        Species Pristimantis simoterus (Lynch, 1980)  
                                        Species Pristimantis siopelus (Lynch and Burrowes, 1990)  
                                        Species Pristimantis sirnigeli Yánez-Muñoz, Meza-Ramos, Cisneros-Heredia and Reyes-Puig, 2011  
                                        Species Pristimantis skydmainos (Flores and Rodríguez, 1997)  
                                        Species Pristimantis sneiderni Ospina-Sarria and Duellman, 2019  
                                        Species Pristimantis sobetes (Lynch, 1980)  
                                        Species Pristimantis spectabilis Duellman and Chaparro, 2008  
                                        Species Pristimantis spilogaster (Lynch, 1984)  
                                        Species Pristimantis spinosus (Lynch, 1979)  
                                        Species Pristimantis stenodiscus (Walker and Test, 1955)  
                                        Species Pristimantis sternothylax (Duellman and Wild, 1993)  
                                        Species Pristimantis stictoboubonus (Duellman, Lehr and Venegas, 2006)  
                                        Species Pristimantis stictogaster (Duellman and Hedges, 2005)  
                                        Species Pristimantis stictus González-Durán, 2016  
                                        Species Pristimantis stipa Venegas and Duellman, 2012  
                                        Species Pristimantis subsigillatus (Boulenger, 1902)  
                                        Species Pristimantis suetus (Lynch and Rueda-Almonacid, 1998)  
                                        Species Pristimantis sulculus (Lynch and Burrowes, 1990)  
                                        Species Pristimantis supernatis (Lynch, 1979)  
                                        Species Pristimantis surdus (Boulenger, 1882)  
                                        Species Pristimantis susaguae (Rueda-Almonacid, Lynch and Galvis-Peñuela, 2003)  
                                        Species Pristimantis taciturnus (Lynch and Suárez-Mayorga, 2003)  
                                        Species Pristimantis taeniatus (Boulenger, 1912)  
                                        Species Pristimantis tamsitti (Cochran and Goin, 1970)  
                                        Species Pristimantis tantanti (Lehr, Torres-Gastello and Suárez-Segovia, 2007)  
                                        Species Pristimantis tanyrhynchus Lehr, 2007  
                                        Species Pristimantis tayrona (Lynch and Ruiz-Carranza, 1985)  
                                        Species Pristimantis telefericus (La Marca, 2005)  
                                        Species Pristimantis tenebrionis (Lynch and Miyata, 1980)  
                                        Species Pristimantis terraebolivaris (Rivero, 1961)  
                                        Species Pristimantis thectopternus (Lynch, 1975)  
                                        Species Pristimantis thyellus (La Marca, 2007)  
                                        Species Pristimantis thymalopsoides (Lynch, 1976)  
                                        Species Pristimantis thymelensis (Lynch, 1972)  
                                        Species Pristimantis tiktik Székely, Eguiguren, Székely, Ordóñez-Delgado, Armijos-Ojeda, Riofrîo-Guamán and Cogalniceanu, 2018  
                                        Species Pristimantis tinajillas Urgilés, Sánchez-Nivicela, Nieves and Yánez-Muñoz, 2014  
                                        Species Pristimantis tinguichaca Brito-M., Ojala-Barbour, Batallas-R. and Almendáriz C., 2016  
                                        Species Pristimantis toftae (Duellman, 1978)  
                                        Species Pristimantis torrenticola (Lynch and Rueda-Almonacid, 1998)  
                                        Species Pristimantis trachyblepharis (Boulenger, 1918)  
                                        Species Pristimantis tribulosus (Lynch and Rueda-Almonacid, 1997)  
                                        Species Pristimantis truebae (Lynch and Duellman, 1997)  
                                        Species Pristimantis tubernasus (Rivero, 1984)  
                                        Species Pristimantis tungurahua Reyes-Puig, Yánez-Muñoz, Cisneros-Heredia and Ramírez, 2011  
                                        Species Pristimantis turik Barrio-Amorós, Rojas-Runjaic and Infante-Rivero, 2008  
                                        Species Pristimantis turpinorum (Hardy, 2001)  
                                        Species Pristimantis turumiquirensis (Rivero, 1961)  
                                        Species Pristimantis uisae (Lynch, 2003)  
                                        Species Pristimantis unistrigatus (Günther, 1859)  
                                        Species Pristimantis urani Rivera-Correa and Daza-R., 2016  
                                        Species Pristimantis uranobates (Lynch, 1991)  
                                        Species Pristimantis urichi (Boettger in Mole and Urich, 1894)  
                                        Species Pristimantis vanadise (La Marca, 1984)  
                                        Species Pristimantis variabilis (Lynch, 1968)  
                                        Species Pristimantis veletis (Lynch and Rueda-Almonacid, 1997)  
                                        Species Pristimantis ventrigranulosus Maciel, Vaz-Silva, Oliveira and Padial, 2012  
                                        Species Pristimantis ventriguttatus Lehr and Köhler, 2007  
                                        Species Pristimantis ventrimarmoratus (Boulenger, 1912)  
                                        Species Pristimantis verecundus (Lynch and Burrowes, 1990)  
                                        Species Pristimantis versicolor (Lynch, 1979)  
                                        Species Pristimantis vertebralis (Boulenger, 1886)  
                                        Species Pristimantis vicarius (Lynch and Ruiz-Carranza, 1983)  
                                        Species Pristimantis vidua (Lynch, 1979)  
                                        Species Pristimantis viejas (Lynch and Rueda-Almonacid, 1999)  
                                        Species Pristimantis vilarsi (Melin, 1941)  
                                        Species Pristimantis vilcabambae Lehr, 2007  
                                        Species Pristimantis vinhai (Bokermann, 1975)  
                                        Species Pristimantis viridicans (Lynch, 1977)  
                                        Species Pristimantis viridis (Ruiz-Carranza, Lynch and Ardila-Robayo, 1997)  
                                        Species Pristimantis w-nigrum (Boettger, 1892)  
                                        Species Pristimantis wagteri (Venegas, 2007)  
                                        Species Pristimantis walkeri (Lynch, 1974)  
                                        Species Pristimantis waoranii (McCracken, Forstner and Dixon, 2007)  
                                        Species Pristimantis wiensi (Duellman and Wild, 1993)  
                                        Species Pristimantis xeniolum (Lynch, 2001)  
                                        Species Pristimantis xestus (Lynch, 1995)  
                                        Species Pristimantis xylochobates (Lynch and Ruiz-Carranza, 1996)  
                                        Species Pristimantis yanezi Navarrete, Venegas and Ron, 2016  
                                        Species Pristimantis yantzaza Valencia, Dueñas, Székely, Batallas-R. and Pulluquitín, 2017  
                                        Species Pristimantis yaviensis (Myers and Donnelly, 1996)  
                                        Species Pristimantis yukpa Barrio-Amorós, Rojas-Runjaic and Infante-Rivero, 2008  
                                        Species Pristimantis yumbo Yánez-Muñoz, Meza-Ramos, Cisneros-Heredia and Reyes-Puig, 2011  
                                        Species Pristimantis yuruaniensis Rödder and Jungfer, 2008  
                                        Species Pristimantis yustizi (Barrio-Amorós and Chacón-Ortiz, 2004)  
                                        Species Pristimantis zeuctotylus (Lynch and Hoogmoed, 1977)  
                                        Species Pristimantis zimmermanae (Heyer and Hardy, 1991)  
                                        Species Pristimantis zoilae (Mueses-Cisneros, 2007)  
                                        Species Pristimantis zophus (Lynch and Ardila-Robayo, 1999)  
       

 References
       
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  Source: Amphibian Species of the World: an Online Reference v6, database (version 6.0)  
  Acquired: 2019   
  Notes: Frost, Darrel R. 2019. Amphibian Species of the World: an Online Reference. Version 6 (12 September, 2019). Electronic Database accessible at http://research.amnh.org/herpetology/amphibia/index.html American Museum of Natural History, New York, USA   
  Reference for: Pristimantis   
       
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Disclaimer: ITIS taxonomy is based on the latest scientific consensus available, and is provided as a general reference source for interested parties. However, it is not a legal authority for statutory or regulatory purposes. While every effort has been made to provide the most reliable and up-to-date information available, ultimate legal requirements with respect to species are contained in provisions of treaties to which the United States is a party, wildlife statutes, regulations, and any applicable notices that have been published in the Federal Register. For further information on U.S. legal requirements with respect to protected taxa, please contact the U.S. Fish and Wildlife Service.

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