ITIS - Report: Exocelina

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Exocelina Broun, 1886
Taxonomic Serial No.: 811209

(Download Help) Exocelina TSN 811209

Taxonomy and Nomenclature


	Kingdom:	Animalia
	Taxonomic Rank:	Genus
	Synonym(s):	Papuadytes Balke, 1998
		Copelatus (Papuadytes) Balke, 1998
	Common Name(s):

	Taxonomic Status:
	Current Standing:	valid

	Data Quality Indicators:
	Record Credibility Rating:	verified - standards met
	Global Species Completeness:	complete
	Latest Record Review:	2021

Taxonomic Hierarchy


	Kingdom	Animalia – Animal, animaux, animals
	Subkingdom	Bilateria – triploblasts
	Infrakingdom	Protostomia
	Superphylum	Ecdysozoa
	Phylum	Arthropoda – Artrópode, arthropodes, arthropods
	Subphylum	Hexapoda – hexapods
	Class	Insecta – insects, hexapoda, inseto, insectes
	Subclass	Pterygota – insects ailés, winged insects
	Infraclass	Neoptera – modern, wing-folding insects
	Superorder	Holometabola
	Order	Coleoptera Linnaeus, 1758 – beetles, coléoptčres, besouro
	Suborder	Adephaga Schellenberg, 1806
	Family	Dytiscidae Leach, 1815 – diving beetles, predaceous diving beetles, ditiques
	Subfamily	Copelatinae Branden, 1885
	Tribe	Copelatini Branden, 1885
	Genus	Exocelina Broun, 1886
		Direct Children:
	Species	Exocelina abdita (Balke, Watts, Cooper, Humphreys and Vogler, 2004)
	Species	Exocelina adelbertensis Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina aipo (Balke, 1998)
	Species	Exocelina aipomek (Balke, 1998)
	Species	Exocelina akameku Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina alexanderi Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina allerbergeri Wewalka, Balke and Hendrich, 2010
	Species	Exocelina amabilis Shaverdo and Balke in Shaverdo et al., 2020
	Species	Exocelina ambua Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina andakombensis Shaverdo and Balke in Shaverdo et al., 2016
	Species	Exocelina anggiensis Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina apistefti Shaverdo, Surbakti and Balke in Shaverdo, Surbakti, Sumoked and Balke, 2021
	Species	Exocelina arfakensis Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina ascendens (Balke, 1998)
	Species	Exocelina aseki Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina astrophallus (Balke, 1998)
	Species	Exocelina athesphati Shaverdo, Surbakti, Sumoked and Balke, 2020
	Species	Exocelina atowaso (Shaverdo, Sagata and Balke, 2005)
	Species	Exocelina atra (Sharp, 1882)
	Species	Exocelina atrata (J. Balfour-Browne, 1939)
	Species	Exocelina atripennis (J. Balfour-Browne, 1939)
	Species	Exocelina aubei (Montrouzier, 1860)
	Species	Exocelina australiae (Clark, 1863)
	Species	Exocelina australis (Clark, 1863)
	Species	Exocelina bacchus Balke in Shaverdo, Surbakti, Sumoked and Balke, 2021
	Species	Exocelina bagus (Balke and Hendrich in Balke, 2001)
	Species	Exocelina baliem Shaverdo, Hendrich and Balke, 2013
	Species	Exocelina barbarae Wewalka, Balke and Hendrich, 2010
	Species	Exocelina bewani Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina bewaniensis Shaverdo, Menufandu and Balke, 2014
	Species	Exocelina bifida Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina bimaculata (Perroud and Montrouzier, 1864)
	Species	Exocelina bismarckensis Shaverdo and Balke, 2014
	Species	Exocelina boulevardi (Watts, 1978)
	Species	Exocelina brahminensis Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina brazza Shaverdo and Balke in Shaverdo et al., 2020
	Species	Exocelina broschii (Balke, 1998)
	Species	Exocelina brownei (Guignot, 1942)
	Species	Exocelina brunoi Wewalka, Balke and Hendrich, 2010
	Species	Exocelina bundiensis Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina burwelli Wewalka, Balke and Hendrich, 2010
	Species	Exocelina casuarina (Balke and Hendrich, 1998)
	Species	Exocelina charlottae Wewalka, Balke and Hendrich, 2010
	Species	Exocelina cheesmanae (J. Balfour-Browne, 1939)
	Species	Exocelina commatifera (Heller, 1916)
	Species	Exocelina craterensis Shaverdo and Balke, 2014
	Species	Exocelina creuxorum Wewalka, Balke and Hendrich, 2010
	Species	Exocelina cyclops Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina damantiensis (Balke, 1998)
	Species	Exocelina danae (Balke, 1998)
	Species	Exocelina desii (Balke, 1999)
	Species	Exocelina edeltraudae Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina ekari Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina ekpliktiki Shaverdo, Surbakti and Balke in Shaverdo, Surbakti, Sumoked and Balke, 2021
	Species	Exocelina elongatula (W. J. MacLeay, 1871)
	Species	Exocelina eme Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina erteldi (Balke, 1998)
	Species	Exocelina evelyncheesmanae Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina ferruginea (Sharp, 1882)
	Species	Exocelina feryi Wewalka, Balke and Hendrich, 2010
	Species	Exocelina flammi Wewalka, Balke and Hendrich, 2010
	Species	Exocelina foja Shaverdo, Surbakti and Balke in Shaverdo, Surbakti, Sumoked and Balke, 2021
	Species	Exocelina fume (Balke, 1998)
	Species	Exocelina gapa (Watts, 1978)
	Species	Exocelina garana Shaverdo and Balke in Shaverdo et al., 2016
	Species	Exocelina gaulorum Wewalka, Balke and Hendrich, 2010
	Species	Exocelina gelima Wewalka, Balke and Hendrich, 2010
	Species	Exocelina glypta (Guignot, 1955)
	Species	Exocelina gorokaensis Shaverdo and Balke, 2014
	Species	Exocelina gracilis (Sharp, 1882)
	Species	Exocelina haia Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina hansferyi Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina heidiae (Balke, 1998)
	Species	Exocelina herowana Shaverdo and Balke, 2014
	Species	Exocelina hintelmannae (Shaverdo, Sagata and Balke, 2005)
	Species	Exocelina hudsoni Shaverdo, Surbakti and Balke in Shaverdo, Surbakti, Sumoked and Balke, 2021
	Species	Exocelina ibalimi Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina inexspectata Wewalka, Balke and Hendrich, 2010
	Species	Exocelina injiensis Shaverdo and Balke in Shaverdo et al., 2016
	Species	Exocelina interrupta (Perroud and Montrouzier, 1864)
	Species	Exocelina iratoi Shaverdo, Wild, Sumoked and Balke, 2017
	Species	Exocelina irianensis Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina jaseminae (Balke, 1998)
	Species	Exocelina jeannae Wewalka, Balke and Hendrich, 2010
	Species	Exocelina jimiensis Shaverdo and Balke, 2014
	Species	Exocelina kabwumensis Shaverdo and Balke in Shaverdo et al., 2016
	Species	Exocelina kailaki Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina kainantuensis (Balke, 2001)
	Species	Exocelina kakapupu Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina karmurensis (Balke, 1998)
	Species	Exocelina keki Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina ketembang (Balke, 1998)
	Species	Exocelina kinibeli Shaverdo and Balke, 2014
	Species	Exocelina kisli Shaverdo and Balke, 2014
	Species	Exocelina knoepfchen Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina kobau Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina koghis Wewalka, Balke and Hendrich, 2010
	Species	Exocelina kolleri Wewalka, Balke and Hendrich, 2010
	Species	Exocelina koroba Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina kowalskii Shaverdo and Balke, 2019
	Species	Exocelina ksionseki Shaverdo and Balke, 2014
	Species	Exocelina kumulensis Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina larsoni (Balke, 1998)
	Species	Exocelina leae Wewalka, Balke and Hendrich, 2010
	Species	Exocelina lembena Shaverdo and Balke, 2014
	Species	Exocelina likui Shaverdo, Wild, Sumoked and Balke, 2017
	Species	Exocelina lilianae Wewalka, Balke and Hendrich, 2010
	Species	Exocelina maculata (Sharp, 1882)
	Species	Exocelina manfredi (Balke, 1998)
	Species	Exocelina manokwariensis Shaverdo, Panjaitan and Balke, 2016
	Species	Exocelina mantembu Shaverdo and Balke, 2014
	Species	Exocelina marawaka Shaverdo and Balke in Shaverdo et al., 2016
	Species	Exocelina marinae (Shaverdo, Sagata and Balke, 2005)
	Species	Exocelina me (Balke, 1998)
	Species	Exocelina mekilensis Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina melanaria (Sharp, 1882)
	Species	Exocelina mendiensis Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina menyamya Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina messeri (Balke, 1999)
	Species	Exocelina mianminensis Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina michaelensis Shaverdo and Balke, 2014
	Species	Exocelina mimika Shaverdo and Balke in Shaverdo et al., 2020
	Species	Exocelina miriae (Balke, 1998)
	Species	Exocelina monae (Balke, 1998)
	Species	Exocelina mondmillensis Shaverdo, Sagata and Balke, 2016
	Species	Exocelina monteithi Wewalka, Balke and Hendrich, 2010
	Species	Exocelina morobensis Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina munaso (Shaverdo, Sagata and Balke, 2005)
	Species	Exocelina nehoue Balke, Hájek, Hendrich and Wewalka, 2014
	Species	Exocelina nielsi Wewalka, Balke and Hendrich, 2010
	Species	Exocelina niklasi Wewalka, Balke and Hendrich, 2010
	Species	Exocelina nomax (J. Balfour-Browne, 1939)
	Species	Exocelina novaecaledoniae (J. Balfour-Browne, 1939)
	Species	Exocelina oceai Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina oiwa Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina okapa Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina okbapensis Shaverdo and Balke in Shaverdo et al., 2017
	Species	Exocelina oksibilensis Shaverdo, Surbakti, Warikar and Balke in Shaverdo et al., 2019
	Species	Exocelina oraia Shaverdo, Surbakti and Balke in Shaverdo, Surbakti, Sumoked and Balke, 2021
	Species	Exocelina ouin Wewalka, Balke and Hendrich, 2010
	Species	Exocelina parvula (Boisduval, 1835)
	Species	Exocelina perfecta (Sharp, 1882)
	Species	Exocelina pinocchi Shaverdo and Balke, 2014
	Species	Exocelina piusi Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina poellabauerae Wewalka, Balke and Hendrich, 2010
	Species	Exocelina polita (Sharp, 1882)
	Species	Exocelina posmani Shaverdo and Balke in Shaverdo et al., 2016
	Species	Exocelina pseudoastrophallus Shaverdo and Balke, 2014
	Species	Exocelina pseudobifida Shaverdo and Balke, 2014
	Species	Exocelina pseudoedeltraudae Shaverdo and Balke, 2014
	Species	Exocelina pseudoeme Shaverdo and Balke, 2014
	Species	Exocelina pseudofume Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina pseudojaseminae Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina pseudomarinae Shaverdo, Sagata and Balke, 2016
	Species	Exocelina pseudopusilla Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina pseudosoppi Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina pui Shaverdo, Wild, Sumoked and Balke, 2017
	Species	Exocelina pulchella Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina pulukensis Shaverdo, Wild, Sumoked and Balke, 2017
	Species	Exocelina punctipennis (Lea, 1899)
	Species	Exocelina pusilla Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina ransikiensis Shaverdo, Panjaitan and Balke, 2016
	Species	Exocelina rasilis (Lea, 1899)
	Species	Exocelina rasjadi Watts and Humphreys, 2009
	Species	Exocelina remyi Wewalka, Balke and Hendrich, 2010
	Species	Exocelina riberai Shaverdo, Surbakti and Balke in Shaverdo, Surbakti, Sumoked and Balke, 2021
	Species	Exocelina rotteri Wewalka, Balke and Hendrich, 2010
	Species	Exocelina rufa (Balke, 1998)
	Species	Exocelina saltusholmesensis Watts, Hendrich and Balke, 2016
	Species	Exocelina saudaunensis Shaverdo and Balke, 2014
	Species	Exocelina schoelleri Wewalka, Balke and Hendrich, 2010
	Species	Exocelina shizong (Balke and Bergsten, 2003)
	Species	Exocelina sima Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina simbaiarea Shaverdo and Balke, 2014
	Species	Exocelina simbaiensis Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina simbaijimi Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina simoni Wewalka, Balke and Hendrich, 2010
	Species	Exocelina simplex (Clark, 1863)
	Species	Exocelina skalei Shaverdo and Balke, 2014
	Species	Exocelina soppi Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina staneki Wewalka, Balke and Hendrich, 2010
	Species	Exocelina subjecta (Sharp, 1882)
	Species	Exocelina sugayai Balke and Ribera, 2020
	Species	Exocelina sumokedi Shaverdo and Balke in Shaverdo et al., 2018
	Species	Exocelina tabubilensis Shaverdo and Balke, 2014
	Species	Exocelina takime (Balke, 1998)
	Species	Exocelina talaki (Balke, 1998)
	Species	Exocelina tariensis Shaverdo and Balke, 2014
	Species	Exocelina tekadu Shaverdo and Balke in Shaverdo et al., 2016
	Species	Exocelina tomhansi Shaverdo, Wild, Sumoked and Balke, 2017
	Species	Exocelina tsinga Shaverdo, Surbakti, Sumoked and Balke, 2020
	Species	Exocelina ullrichi (Balke, 1998)
	Species	Exocelina unipo Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina utowaensis Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina varirata Shaverdo and Balke in Shaverdo et al., 2016
	Species	Exocelina vladimiri (Shaverdo, Sagata and Balke, 2005)
	Species	Exocelina vovai Shaverdo and Balke, 2014
	Species	Exocelina waaf Shaverdo, Surbakti and Balke in Shaverdo, Surbakti, Sumoked and Balke, 2021
	Species	Exocelina waigeoensis Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina wannangensis Shaverdo and Balke, 2014
	Species	Exocelina warahulenensis Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina warasera Shaverdo and Balke in Shaverdo et al., 2019
	Species	Exocelina wareaga Shaverdo and Balke in Shaverdo et al., 2016
	Species	Exocelina weylandensis Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina wigodukensis Shaverdo, Wild, Sumoked and Balke, 2017
	Species	Exocelina woitapensis Shaverdo and Balke in Shaverdo et al., 2016
	Species	Exocelina wondiwoiensis Shaverdo, Hendrich and Balke, 2012
	Species	Exocelina yoginofi Shaverdo and Balke in Shaverdo et al., 2018

References


	Expert(s):
	Expert:
	Notes:
	Reference for:

	Other Source(s):
	Source:	A World Catalogue of the Family Dytiscidae (2021), database (version 1.I.2021)
	Acquired:	2021
	Notes:	Nilsson A.N. & Hájek J. 2021: A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1.I.2021, 315 pp. URL: http://www.waterbeetles.eu/ Accessed 1 Jul 2021
	Reference for:	Exocelina

	Publication(s):
	Author(s)/Editor(s):	Watts, C. H. S., and W. F. Humphreys
	Publication Date:	2009
	Article/Chapter Title:	Fourteen new Dytiscidae (Coleoptera) of the genera Limbodessus Guignot, Paroster Sharp, and Exocelina Broun from underground waters in Australia
	Journal/Book Name, Vol. No.:	Transactions of the Royal Society of South Australia, vol. 133, no. 1
	Page(s):	62-107
	Publisher:
	Publication Place:
	ISBN/ISSN:	0372-1426
	Notes:
	Reference for:	Exocelina

Geographic Information


	Geographic Division:

	Jurisdiction/Origin:

Comments


	Comment:

Disclaimer: ITIS taxonomy is based on the latest scientific consensus available, and is provided as a general reference source for interested parties. However, it is not a legal authority for statutory or regulatory purposes. While every effort has been made to provide the most reliable and up-to-date information available, ultimate legal requirements with respect to species are contained in provisions of treaties to which the United States is a party, wildlife statutes, regulations, and any applicable notices that have been published in the Federal Register. For further information on U.S. legal requirements with respect to protected taxa, please contact the U.S. Fish and Wildlife Service.

Integrated Taxonomic Information System - Report