Frost, Darrel R. 2019. Amphibian Species of the World: an Online Reference. Version 6 (12 September, 2019). Electronic Database accessible at http://research.amnh.org/herpetology/amphibia/index.html American Museum of Natural History, New York, USA
A Solution to the Large Black Salamander Problem (Genus Bolitoglossa) in Costa Rica and Panamá
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Copeia, vol. 2005, no. 2
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