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Heriades  Spinola, 1808
Taxonomic Serial No.: 154375

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 Taxonomy and Nomenclature
  Kingdom: Animalia  
  Taxonomic Rank: Genus  
  Common Name(s):    
  Taxonomic Status:    
  Current Standing: valid  
  Data Quality Indicators:    
  Record Credibility Rating: verified - standards met  
  Global Species Completeness: complete   
  Latest Record Review: 2008   

 Taxonomic Hierarchy
 KingdomAnimalia  – Animal, animaux, animals  
             PhylumArthropoda  – Artrópode, arthropodes, arthropods  
                SubphylumHexapoda  – hexapods  
                   ClassInsecta  – insects, hexapoda, inseto, insectes  
                      SubclassPterygota  – insects ailés, winged insects  
                         InfraclassNeoptera  – modern, wing-folding insects  
                               OrderHymenoptera  – abelha, formiga, vespa, ants, bees, wasps  
                                  SuborderApocrita  – abeilles, fourmis, guępes véritables, narrow-waisted hymenopterans, ants, bees, true wasps  
                                        SuperfamilyApoidea  – bees, sphecoid wasps, apoid wasps  
                                           FamilyMegachilidae  – leafcutting bees  
                                                    GenusHeriades Spinola, 1808  
    Direct Children:  
                                                       Species Heriades abessinicus Friese, 1915  
                                                       Species Heriades albiscopanus Strand, 1912  
                                                       Species Heriades aldabranus Cockerell, 1912  
                                                       Species Heriades alfkeni Benoist, 1938  
                                                       Species Heriades ambiguus Cockerell, 1946  
                                                       Species Heriades ammodendri Popov, 1960  
                                                       Species Heriades angusticeps Cockerell, 1937  
                                                       Species Heriades angustulus Cockerell, 1937  
                                                       Species Heriades apriculus Griswold, 1998  
                                                       Species Heriades arcuatellus Cockerell, 1937  
                                                       Species Heriades arnoldi Friese, 1922  
                                                       Species Heriades bequaerti Cockerell, 1931  
                                                       Species Heriades binghami Dover, 1925  
                                                       Species Heriades binodosus Cockerell, 1936  
                                                       Species Heriades bouyssoui Vachal, 1903  
                                                       Species Heriades brage Strand, 1912  
                                                       Species Heriades bruneri Titus, 1904  
                                                       Species Heriades burgeoni Benoist, 1931  
                                                       Species Heriades canaliculatus Benoist, 1931  
                                                       Species Heriades cancavus Wu, 1982  
                                                       Species Heriades capicola Strand, 1912  
                                                       Species Heriades carinata Cresson, 1864  
                                                       Species Heriades chariensis Benoist, 1931  
                                                       Species Heriades chrysogaster (Cameron, 1897)  
                                                       Species Heriades chubbi Cockerell, 1916  
                                                       Species Heriades cingulata Benoist, 1931  
                                                       Species Heriades civicus Cockerell, 1937  
                                                       Species Heriades clavicornis Morawitz, 1875  
                                                       Species Heriades crenulatus Nylander, 1856  
                                                       Species Heriades cressoni Michener, 1938  
                                                       Species Heriades crucifer Cockerell, 1897  
                                                       Species Heriades currani Michener, 1943  
                                                       Species Heriades curviventris Strand, 1912  
                                                       Species Heriades dalmaticus Maidl, 1922  
                                                       Species Heriades debilicornis Cockerell, 1940  
                                                       Species Heriades diminutus Cockerell, 1937  
                                                       Species Heriades discrepans Benoist, 1938  
                                                       Species Heriades ellenbergeri Benoist, 1931  
                                                       Species Heriades erythrosoma Friese, 1922  
                                                       Species Heriades eximius Friese, 1904  
                                                       Species Heriades fertoni Benoist, 1938  
                                                       Species Heriades filicornis Friese, 1915  
                                                       Species Heriades flocciferus Brauns, 1929  
                                                       Species Heriades freygessneri Schletterer, 1889  
                                                       Species Heriades frontosus Schletterer, 1889  
                                                       Species Heriades fuelleborni Friese, 1922  
                                                       Species Heriades fujiyamai Yasumatsu & Hirashima, 1952  
                                                       Species Heriades fulleborni Friese, 1922  
                                                       Species Heriades fulvescens Cockerell, 1920  
                                                       Species Heriades fulvohispidus Yasumatsu & Hirashima, 1952  
                                                       Species Heriades gibbosus (Friese, 1909)  
                                                       Species Heriades gracilior Cockerell, 1897  
                                                       Species Heriades hercules Strand, 1911  
                                                       Species Heriades hierosolomitus Benoist, 1935  
                                                       Species Heriades hissaricus Popov, 1955  
                                                       Species Heriades impressus Schletterer, 1889  
                                                       Species Heriades ingogoensis Cockerell, 1946  
                                                       Species Heriades labiatus Pérez, 1896  
                                                       Species Heriades langenburgicus Strand, 1911  
                                                       Species Heriades laosellus Cockerell, 1929  
                                                       Species Heriades latipes Popov, 1960  
                                                       Species Heriades leavitti Crawford, 1913  
                                                       Species Heriades libericus Cockerell, 1931  
                                                       Species Heriades longicornis (Friese, 1915)  
                                                       Species Heriades longicornis Popov, 1960  
                                                       Species Heriades longispinis Cockerell, 1944  
                                                       Species Heriades macrognatus Vachal, 1909  
                                                       Species Heriades mamilliferus Brauns, 1929  
                                                       Species Heriades mandibularis Friese, 1922  
                                                       Species Heriades micheneri Timberlake, 1947  
                                                       Species Heriades micropthalma Michener, 1954  
                                                       Species Heriades microstictus Cockerell, 1936  
                                                       Species Heriades mundulus Cockerell, 1920  
                                                       Species Heriades nasiferus Cockerell, 1946  
                                                       Species Heriades nitescens Cockerell, 1931  
                                                       Species Heriades nodulosus Cockerell, 1937  
                                                       Species Heriades occidentalis Michener, 1938  
                                                       Species Heriades orientalis Gupta, 1987  
                                                       Species Heriades ornaticornis Cockerell, 1936  
                                                       Species Heriades otavica Eardley & R. P. Urban, 2006  
                                                       Species Heriades othonis Friese, 1914  
                                                       Species Heriades pachyacanthus Cockerell, 1940  
                                                       Species Heriades paganensis Yasumatsu, 1942  
                                                       Species Heriades parvulus Bingham, 1897  
                                                       Species Heriades patellus (Nurse, 1902)  
                                                       Species Heriades perminutus Cockerell, 1935  
                                                       Species Heriades perpolitus Cockerell, 1946  
                                                       Species Heriades phthisica Gerstäcker, 1858  
                                                       Species Heriades plumosus Krombein, 1950  
                                                       Species Heriades pogonura Benoist, 1931  
                                                       Species Heriades pretorii Cockerell, 1946  
                                                       Species Heriades prionsa (Cameron, 1905)  
                                                       Species Heriades psiadiae Pauly & Griswold, 2001  
                                                       Species Heriades punctulatus Cockerell, 1917  
                                                       Species Heriades punctulifera Schletterer, 1889  
                                                       Species Heriades rowlandi (Cockerell, 1947)  
                                                       Species Heriades rubicolus Pérez, 1890  
                                                       Species Heriades rufifrons Cockerell, 1932  
                                                       Species Heriades sakishimanus Yasumatsu & Hirashima, 1965  
                                                       Species Heriades sauteri Cockerell, 1911  
                                                       Species Heriades schwarzi Griswold, 1998  
                                                       Species Heriades scutellatus Friese, 1922  
                                                       Species Heriades seyrigi Pauly & Griswold, 2001  
                                                       Species Heriades shestakovi Popov, 1960  
                                                       Species Heriades sinuatus Spinola, 1808  
                                                       Species Heriades spiniscutis (Cameron, 1905)  
                                                       Species Heriades strictifrons Cockerell, 1946  
                                                       Species Heriades subfrontosus Cockerell, 1939  
                                                       Species Heriades sulcatiferus Cockerell, 1937  
                                                       Species Heriades sulcatifrons Cockerell, 1936  
                                                       Species Heriades sulcatulus Cockerell, 1931  
                                                       Species Heriades tenuis Nurse, 1904  
                                                       Species Heriades testaceicornis (Cameron, 1908)  
                                                       Species Heriades texanus Michener, 1938  
                                                       Species Heriades timberlakei Michener, 1938  
                                                       Species Heriades trigibbiferus Brauns, 1929  
                                                       Species Heriades truncorum (Linnaeus, 1758)  
                                                       Species Heriades turcomanicus Popov, 1955  
                                                       Species Heriades usakensis Cockerell, 1937  
                                                       Species Heriades ustulata Benoist, 1931  
                                                       Species Heriades variolosus (Cresson, 1872)  
                                                       Species Heriades victoriae Friese, 1922  
                                                       Species Heriades wellmani Cockerell, 1908  
                                                       Species Heriades wilmattae Cockerell, 1931  
                                                       Species Heriades xanthogaster Cockerell, 1932  
                                                       Species Heriades yunnanensis Wu, 1992  

  Reference for:    
  Other Source(s):    
  Source: Catalogue of Bees (Hymenoptera, Apoidea) in the Neotropical..., website (version 03July2012)  
  Acquired: 2016   
  Notes: Moure, J. S., Urban, D., and G. A. R. Melo (Orgs). Catalogue of Bees (Hymenoptera, Apoidea) in the Neotropical Region. Available online at   
  Reference for: Heriades   
  Source: Hymenoptera Name Server version 0.03 4.x.2002, website (version 0.03)  
  Acquired: 2004   
  Reference for: Heriades   
  Source: NODC Taxonomic Code, database (version 8.0)  
  Acquired: 1996   
  Reference for: Heriades   
  Author(s)/Editor(s): Michener, Charles D.  
  Publication Date: 2000   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: The Bees of the World   
  Page(s): xiv + 913   
  Publisher: The Johns Hopkins University Press   
  Publication Place: Baltimore, Maryland, USA   
  ISBN/ISSN: 0-8018-6133-0   
  Reference for: Heriades   
  Author(s)/Editor(s): Michener, Charles D.  
  Publication Date: 2007   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: The Bees of the World, Second Edition   
  Page(s): xvi + 953   
  Publisher: The Johns Hopkins University Press   
  Publication Place: Baltimore, Maryland, USA   
  ISBN/ISSN: 0-8018-8573-6   
  Reference for: Heriades   
  Author(s)/Editor(s): Spinola, M.  
  Publication Date: 1808   
  Article/Chapter Title:    
  Journal/Book Name, Vol. No.: Insectorum Liguriae Species Novae aut Rariores, quas in agro Ligustico nuper detexit, descripsit, et iconibus illustravit Maximilianus Spinola, adjecto Catalogo spiecierum auctoribus jam enumeratarum, quae in eadam regione occurrunt, vol. 2   
  Page(s): ii + 262   
  Publisher: Gravier   
  Publication Place: Genuae (Genoa), Italy   
  Notes: Available online via Biodiversity Heritage Library at   
  Reference for: original description   
  Author(s)/Editor(s): Ungricht, Stefan, Andreas Müller, and Silvia Dorn  
  Publication Date: 2008   
  Article/Chapter Title: A taxonomic catalogue of the Palaearctic bees of the tribe Osmiini (Hymenoptera: Apoidea: Megachilidae)   
  Journal/Book Name, Vol. No.: Zootaxa, issue 1865   
  Page(s): 253   
  Publication Place:    
  ISBN/ISSN: 9781869772550, 1175-5326  
  Notes: Online edition ISBN is 978-1-86977-256-7 and ISSN is 1175-5334:   
  Reference for: Heriades   

 Geographic Information
  Geographic Division:    



  Comment: The gender of Heriades Spinola 1808 is feminine. Spinola (1808: 7) proposed Heriades, and included several species without giving explicit combinations in the new genus. However, he did (p.9) present two new species names, as 'Heriades pusilla, [H.] sinuata, mihi,' which clearly indicates his use of the genus name as feminine. An etymological commentary by Doug Yanega (Yahoo BeeMonitoring Listserv, 5 October, 2015) is here noted, for reference: 'Dalla Torre and Friese were absolutely wrong; there is no form of the noun for wool - either "eiros" or "erion" - which would be spelled as "Eriades". The term "eriodes" would mean "woolly". If you look at Spinola, his entire paper was written in Latin, and he uses the forms "Heriadibus" and "Heriadum", which indicate that the word "Heriades" is a 3rd-declension plural noun, so the singular form would be "Heriad" - either a son of Eris (which would be masculine), or a mountain nymph (which would be feminine). He included two feminine species, so we know which of the two he intended'  



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