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Publication: McEachran, John D., and Katherine A. Dunn. 1998. Phylogenetic Analysis of Skates, a Morphologically Conservative Clade of Elasmobranchs (Chondrichthyes: Rajidae). Copeia, vol. 1998, no. 2. 271-290.

ITIS records citing this publication
 
 Kingdom Animalia
  Amblyraja Malm, 1877 -- valid -- stout skates
  Amblyraja badia (Garman, 1899) -- valid
  Amblyraja doellojuradoi (Pozzi, 1935) -- valid
  Amblyraja frerichsi Krefft, 1968 -- valid
  Amblyraja georgiana (Norman, 1938) -- valid
  Amblyraja hyperborea (Collett, 1879) -- valid -- Arctic skate, Artic skate
  Amblyraja jenseni (Bigelow and Schroeder, 1950) -- valid -- shorttail skate
  Amblyraja radiata (Donovan, 1808) -- valid -- raie épineuse, starry skate, thorny skate
  Amblyraja reversa (Lloyd, 1906) -- valid
  Amblyraja robertsi (Hulley, 1970) -- valid
  Amblyraja taaf (Meissner, 1987) -- valid
  Amblyrajini -- invalid -- stout skates
  Anacanthobatis von Bonde and Swart, 1923 -- valid -- smooth leg skate
  Anacanthobatis americanus Bigelow and Schroeder, 1962 -- valid -- American leg skate
  Anacanthobatis borneensis Chan, 1965 -- valid
  Anacanthobatis donghaiensis (Deng, Xiong and Zhan, 1983) -- valid
  Anacanthobatis folirostris (Bigelow and Schroeder, 1951) -- valid -- leaf-nose leg skate
  Anacanthobatis longirostris Bigelow and Schroeder, 1962 -- valid -- longnose leg skate
  Anacanthobatis marmoratus von Bonde and Swart, 1923 -- valid
  Anacanthobatis melanosoma (Chan, 1965) -- valid
  Anacanthobatis nanhaiensis (Meng and Li, in Chu, Meng, Hu and Li, 1981) -- valid
  Anacanthobatis ori (Wallace, 1967) -- valid
  Anacanthobatis stenosomus (Li and Hu in Chu, Meng, Hu and Li, 1982) -- valid
  Arhynchobatinae -- invalid
  Arhynchobatini -- invalid
  Arhynchobatis Waite, 1909 -- valid
  Arhynchobatis asperrimus Waite, 1909 -- valid
  Atlantoraja Menni, 1972 -- valid
  Atlantoraja castelnaui (Miranda-Ribeiro, 1907) -- valid
  Atlantoraja cyclophora (Regan, 1903) -- valid
  Atlantoraja platana (Günther, 1880) -- valid
  Bathyraja Ishiyama, 1958 -- valid
  Bathyraja abyssicola (Gilbert, 1896) -- valid -- deepsea skate
  Bathyraja aguja (Kendall and Radcliffe, 1912) -- valid
  Bathyraja albomaculata (Norman, 1937) -- valid
  Bathyraja aleutica (Gilbert, 1896) -- valid -- Aleutian skate
  Bathyraja andriashevi Dolganov, 1985 -- valid
  Bathyraja bergi Dolganov, 1985 -- valid
  Bathyraja brachyurops (Fowler, 1910) -- valid
  Bathyraja caeluronigricans Ishiyama and Ishihara, 1977 -- valid
  Bathyraja diplotaena (Ishiyama, 1952) -- valid
  Bathyraja eatonii (Günther, 1876) -- valid
  Bathyraja federovi Dolganov, 1985 -- valid
  Bathyraja griseocauda (Norman, 1937) -- valid
  Bathyraja hesperafricana Stehmann, 1995 -- valid
  Bathyraja hubbsi Ishihara and Ishiyama, 1985 -- valid -- mud skate
  Bathyraja interrupta (Gill and Townsend, 1897) -- valid -- Bering skate, sandpaper skate
  Bathyraja irrasa Hureau and Ozouf-Costaz, 1980 -- valid
  Bathyraja isotrachys (Günther, 1877) -- valid
  Bathyraja lindbergi Ishiyama and Ishihara, 1977 -- valid -- Commander skate
  Bathyraja longicauda (de Buen, 1959) -- valid
  Bathyraja maccaini Springer, 1971 -- valid
  Bathyraja macloviana (Norman, 1937) -- valid
  Bathyraja maculata Ishiyama and Ishihara, 1977 -- valid -- whiteblotched skate
  Bathyraja magellanica (Philippi, 1902) -- valid
  Bathyraja matsubarai (Ishiyama, 1952) -- valid
  Bathyraja meridionalis Stehmann, 1987 -- valid
  Bathyraja minispinosa Ishiyama and Ishihara, 1977 -- valid
  Bathyraja multispinis (Norman, 1937) -- valid
  Bathyraja murrayi (Günther, 1880) -- valid
  Bathyraja notoroensis Ishiyama and Ishihara, 1977 -- valid
  Bathyraja pallida (Forster, 1967) -- valid
  Bathyraja papilionifera Stehmann, 1985 -- valid
  Bathyraja parmifera (Bean, 1881) -- valid -- Alaska skate
  Bathyraja peruana McEachran and Miyake, 1984 -- valid
  Bathyraja pseudoisotrachys Ishihara and Ishiyama, 1985 -- valid
  Bathyraja richardsoni (Garrick, 1961) -- valid -- deepwater skate
  Bathyraja scaphiops (Norman, 1937) -- valid -- cuphead skate
  Bathyraja schroederi (Krefft, 1968) -- valid
  Bathyraja shuntovi Dolganov, 1985 -- valid
  Bathyraja simoterus (Ishiyama, 1967) -- valid
  Bathyraja smirnovi (Soldatov and Pavlenko, 1915) -- valid
  Bathyraja smithii (Müller and Henle, 1841) -- valid
  Bathyraja spinicauda (Jensen, 1914) -- valid -- spinytail skate
  Bathyraja spinosissima (Beebe and Tee-van, 1941) -- valid
  Bathyraja trachouros (Ishiyama, 1958) -- valid
  Bathyraja trachura (Gilbert, 1892) -- valid -- roughtail skate
  Bathyraja tzinovskii Dolganov, 1985 -- valid
  Bathyraja violacea (Suvorov, 1935) -- valid -- Okhotsk skate
  Breviraja Bigelow and Schroeder, 1948 -- valid -- lightnose skates
  Breviraja claramaculata McEachran and Matheson, 1985 -- valid -- brightspot skate
  Breviraja colesi Bigelow and Schroeder, 1948 -- valid -- lightnose skate
  Breviraja cubensis Bigelow and Schroeder, 1950 -- invalid
  Breviraja ishiyamai Bigelow and Schroeder, 1962 -- invalid
  Breviraja marklei McEachran and Miyake, 1987 -- valid -- Markle's skate
  Breviraja mouldi McEachran and Matheson, 1995 -- valid -- blacknose skate
  Breviraja nigriventralis McEachran and Matheson, 1985 -- valid -- blackbelly skate
  Breviraja pallida Forster, 1967 -- invalid
  Breviraja plutonia Garman, 1881 -- invalid -- longtail skate
  Breviraja sinusmexicanus (Bigelow and Schroeder, 1950) -- invalid
  Breviraja spinacidermis (Barnard, 1923) -- invalid
  Breviraja spinosa Bigelow and Schroeder, 1950 -- valid -- spinose skate
  Breviraja yucatanensis Bigelow and Schroeder, 1950 -- invalid
  Cruriraja Bigelow and Schroeder, 1948 -- valid -- rough leg skates
  Cruriraja andamanica (Lloyd, 1909) -- valid
  Cruriraja atlantis Bigelow and Schroeder, 1948 -- valid -- Atlantic leg skate, Atlantic legskate
  Cruriraja cadenati Bigelow and Schroeder, 1962 -- valid -- broadfoot leg skate
  Cruriraja parcomaculata (Von Bonde and Swart, 1923) -- valid
  Cruriraja poeyi Bigelow and Schroeder, 1948 -- valid -- Cuban leg skate, Cuban legskate
  Cruriraja rugosa Bigelow and Schroeder, 1958 -- valid -- rough leg skate
  Cruriraja triangularis Smith, 1964 -- valid
  Dactylobatus Bean and Weed, 1909 -- valid -- skilletskates
  Dactylobatus armatus Bean and Weed, 1909 -- valid -- skilletskate
  Dactylobatus clarkii (Bigelow and Schroeder, 1958) -- valid
  Dipturus Rafinesque, 1810 -- valid -- longnosed skates
  Dipturus batis (Linnaeus, 1758) -- valid -- blue skate, flapper skate
  Dipturus bullisi (Bigelow and Schroeder, 1962) -- valid -- lozenge skate, raya triangular, Tortugas skate
  Dipturus campbelli (Wallace, 1967) -- valid
  Dipturus chilensis (Guichenot, 1848) -- valid
  Dipturus crosnieri (Seret, 1989) -- valid
  Dipturus doutrei (Cadenat, 1960) -- valid
  Dipturus ecuadoriensis (Beebe and Tee-van, 1941) -- valid
  Dipturus garricki (Bigelow and Schroeder, 1958) -- valid -- San Blas skate
  Dipturus gigas (Ishiyama, 1958) -- valid
  Dipturus gudgeri (Whitley, 1940) -- valid
  Dipturus innominatus (Garrick and Paul, 1974) -- valid
  Dipturus johannisdavisi (Alcock, 1899) -- valid
  Dipturus kwangtungensis (Chu, 1960) -- valid
  Dipturus laevis (Mitchill, 1818) -- valid -- barndoor skate, grande raie
  Dipturus lanceorostratus (Wallace, 1967) -- valid
  Dipturus leptocauda (Krefft and Stehmann, 1975) -- valid
  Dipturus linteus (Fries, 1838) -- valid
  Dipturus macrocauda (Ishiyama, 1955) -- valid
  Dipturus nasutus (Müller and Henle, 1841) -- valid
  Dipturus nidarosiensis (Storm, 1881) -- valid
  Dipturus nidrosiensis (Storm, 1881) -- invalid
  Dipturus olseni (Bigelow and Schroeder, 1951) -- valid -- raya colona, spreadfin skate
  Dipturus oregoni (Bigelow and Schroeder, 1958) -- valid -- hooktail skate
  Dipturus oxyrinchus (Linnaeus, 1758) -- valid -- longnosed skate
  Dipturus pullopunctatus (Smith, 1964) -- valid
  Dipturus springeri (Wallace, 1967) -- valid
  Dipturus stenorhynchus (Wallace, 1967) -- valid
  Dipturus teevani (Bigelow and Schroeder, 1951) -- valid -- Caribbean skate
  Dipturus tengu (Jordan and Fowler, 1903) -- valid
  Dipturus trachyderma (Krefft and Stehmann, 1975) -- valid
  Fenestraja McEachran and Compagno, 1982 -- valid -- pygmy skates
  Fenestraja atripinna (Bigelow and Schroeder, 1950) -- valid -- blackfin pygmy skate, blackfin skate, raya aletinegra
  Fenestraja cubensis (Bigelow and Schroeder, 1950) -- valid -- Cuban pygmy skate
  Fenestraja ishiyamai (Bigelow and Schroeder, 1962) -- valid -- plain pygmy skate
  Fenestraja maceachrani (Seret, 1989) -- valid
  Fenestraja mamillidens (Alcock, 1889) -- valid
  Fenestraja plutonia (Garman, 1881) -- valid -- longtail skate, Pluto skate
  Fenestraja sibogae (Weber, 1913) -- valid
  Fenestraja sinusmexicanus (Bigelow and Schroeder, 1950) -- valid -- Gulf of Mexico pygmy skate, Gulf skate, raya pigmea
  Gurgesiella de Buen, 1959 -- valid -- Finless pygmy skates
  Gurgesiella atlantica (Bigelow and Schroeder, 1962) -- valid
  Gurgesiella dorsalifera McEachran and Campagno, 1980 -- valid
  Gurgesiella furvescens de Buen, 1959 -- valid
  Gurgesiellini -- invalid
  Irolita Whitley, 1931 -- valid -- round skates
  Irolita waitii (McCulloch, 1911) -- valid
  Leucoraja Malm, 1877 -- valid -- rough skates
  Leucoraja circularis (Couch, 1838) -- valid -- cuckoo ray, sandy ray
  Leucoraja compagnoi (Stehmann, 1995) -- valid
  Leucoraja erinacea (Mitchill, 1825) -- valid -- common skate, little skate, raie-hérisson, summer skate
  Leucoraja fullonica (Linnaeus, 1758) -- valid -- shagreen ray, shagreen skate
  Leucoraja garmani (Whitley, 1939) -- valid -- rosette skate
  Leucoraja lentiginosa (Bigelow and Schroeder, 1951) -- valid -- freckled skate, raya pecosa, speckled skate
  Leucoraja leucosticta (Stehmann, 1971) -- valid
  Leucoraja melitensis (Clark, 1926) -- valid
  Leucoraja naevus (Müller and Henle, 1841) -- valid -- cuckoo ray, sandy ray
  Leucoraja ocellata (Mitchill, 1815) -- valid -- big skate, eyed skate, raie tachetée, winter skate
  Leucoraja wallacei (Hulley, 1970) -- valid
  Leucoraja yucatanensis (Bigelow and Schroeder, 1950) -- valid -- Yucatan skate
  Malacoraja Stehmann, 1970 -- valid -- soft skates
  Malacoraja kreffti Stehmann, 1977 -- valid
  Malacoraja senta (Garman, 1885) -- valid -- raie à queue de velours, smooth skate
  Malacoraja spinacidermis (Barnard, 1923) -- valid -- soft skate
  Neoraja McEachran and Campagno, 1982 -- valid -- pygmy skates
  Neoraja africana (Stehmann and Seret, 1983) -- valid
  Neoraja caerulea (Stehmann, 1976) -- valid
  Neoraja carolinensis McEachran and Stehmann, 1984 -- valid -- Carolina pygmy skate
  Neoraja stehmanni (Hulley, 1972) -- valid
  Notoraja Ishiyama, 1958 -- valid -- velvet skates
  Notoraja asperula (Garrick and Paul, 1974) -- valid
  Notoraja laxipella (Yearsley and Last, 1992) -- valid
  Notoraja ochroderma McEachran and Last, 1994 -- valid
  Notoraja spinifera (Garrick and Paul, 1974) -- valid
  Notoraja subtilispinosa Stehmann, 1989 -- valid
  Notoraja tobitukai (Hiyama, 1940) -- valid
  Okamejei Ishiyama, 1958 -- valid -- spiny rasp skates
  Okamejei acutispina (Ishiyama, 1958) -- valid
  Okamejei australis (Macleay, 1884) -- valid
  Okamejei boesemani (Ishihara, 1987) -- valid
  Okamejei cerva (Whitley, 1939) -- valid
  Okamejei heemstrai (McEachran and Fechhelm, 1982) -- valid
  Okamejei hollandi (Jordan and Richardson, 1909) -- valid
  Okamejei kenojei (Müller and Henle, 1841) -- valid
  Okamejei lemprieri (Richardson, 1845) -- valid
  Okamejei meerdervoortii (Bleeker, 1860) -- valid
  Okamejei pita (Fricke and Al-Hussar, 1995) -- valid
  Okamejei powelli (Alcock, 1898) -- valid
  Okamejei schmidti (Ishiyama, 1958) -- valid
  Pavoraja Whitley, 1939 -- valid -- peacock skates
  Pavoraja alleni McEachran and Fechhelm, 1982 -- valid
  Pavoraja nitida (Günther, 1880) -- valid
  Psammobatis Günther, 1870 -- valid
  Psammobatis bergi Marini, 1932 -- valid
  Psammobatis extenta (Garman, 1913) -- valid
  Psammobatis lentiginosa McEachran, 1983 -- valid
  Psammobatis normani McEachran, 1983 -- valid
  Psammobatis parvacauda McEachran, 1983 -- valid
  Psammobatis rudis Günther, 1870 -- valid
  Psammobatis rutrum Jordan, 1890 -- valid
  Psammobatis scobina (Philippi, 1857) -- valid
  Pseudoraja Bigelow and Schroeder, 1954 -- valid
  Pseudoraja fischeri Bigelow and Schroeder, 1954 -- valid -- fanfin skate
  Pseudoraja fisheri Bigelow and Schroeder, 1954 -- invalid
  Raja ackleyi Garman, 1881 -- valid -- ocellate skate, raya ocelada
  Raja africana Capapé, 1977 -- valid
  Raja asterias Delaroche, 1809 -- valid -- starry skate
  Raja bahamensis Bigelow and Schroeder, 1965 -- valid -- Bahama skate
  Raja bathyphilia Holt and Byrne, 1908 -- invalid -- abyssal skate
  Raja batis Linnaeus, 1758 -- invalid -- blue skate, flapper skate
  Raja binoculata Girard, 1855 -- valid -- big skate, raya bruja gigante
  Raja brachyura Lafont, 1873 -- valid -- blonde ray
  Raja cervigoni Bigelow and Schroeder, 1964 -- valid -- Venezuela skate
  Raja clavata Linnaeus, 1758 -- valid -- thornback ray, thornback skate
  Raja cortezensis McEachran and Miyake, 1988 -- valid -- Cortez skate, raya de Cortés
  Raja eglanteria Bosc in Lacepède, 1800 -- valid -- clearnose skate, raya naricita
  Raja equatorialis Jordan and Bollman, 1890 -- valid -- equatorial skate, raya ecuatorial
  Raja floridana Bigelow and Schroeder, 1962 -- invalid
  Raja herwigi Krefft, 1965 -- valid
  Raja inornata Jordan and Gilbert, 1881 -- valid -- California skate, raya de California
  Raja maderensis Lowe, 1838 -- valid
  Raja microocellata Montagu, 1818 -- valid -- painted ray
  Raja miraletus Linnaeus, 1758 -- valid -- brown ray, twineye skate
  Raja montagui Fowler, 1910 -- valid -- homelyn ray, spotted ray
  Raja polystigma Regan, 1923 -- valid
  Raja pulchra Liu, 1932 -- valid
  Raja radula Delaroche, 1809 -- valid
  Raja rhina Jordan and Gilbert, 1880 -- valid -- longnose skate, raya narigona
  Raja rosispinis Gill and Townsend, 1897 -- invalid -- flathead skate
  Raja stellulata Jordan and Gilbert, 1880 -- valid -- raya estrellada, starry skate
  Raja straeleni Poll, 1951 -- valid
  Raja texana Chandler, 1921 -- valid -- raya tigre, roundel skate
  Raja undulata Lacepede, 1802 -- valid -- painted ray, undulate ray
  Raja velezi Chirichigno F., 1973 -- valid -- rasptail skate, raya chillona
  Rajella Stehmann, 1970 -- valid -- gray skates
  Rajella annandalei (Weber, 1913) -- valid
  Rajella barnardi (Norman, 1935) -- valid
  Rajella bathyphila (Holt and Byrne, 1908) -- valid -- abyssal skate, deepwater ray, deepwater skate
  Rajella bigelowi (Stehmann, 1978) -- valid -- Bigelow's ray, Bigelow's skate
  Rajella caudaspinosa (Von Bonde and Swart, 1923) -- valid
  Rajella dissimilis (Hulley, 1970) -- valid
  Rajella fuliginea (Bigelow and Schroeder, 1954) -- valid -- sooty skate
  Rajella fyllae (Lütken, 1887) -- valid -- round skate
  Rajella kukujevi (Dolganov, 1985) -- valid
  Rajella leopardus (Von Bonde and Swart, 1923) -- valid
  Rajella nigerrima (de Buen, 1960) -- valid
  Rajella purpuriventralis (Bigelow and Schroeder, 1962) -- valid -- purplebelly skate
  Rajella ravidula (Hulley, 1970) -- valid
  Rajella sadowskii (Krefft and Stehmann, 1974) -- valid
  Rajinae -- invalid
  Rajini -- invalid
  Rhinoraja Ishiyama, 1952 -- valid -- jointnose skates
  Rhinoraja kujiensis (Tanaka, 1916) -- valid
  Rhinoraja longi Raschi and McEachran, 1991 -- valid
  Rhinoraja longicauda Ishiyama, 1952 -- valid
  Rhinoraja odai Ishiyama, 1958 -- valid
  Rhinoraja taranetzi Dolganov, 1985 -- valid
  Rioraja Whitley, 1939 -- valid -- Rio skates
  Rioraja agassizii (Müller and Henle, 1841) -- valid
  Riorajini -- invalid
  Rostroraja Hulley, 1972 -- valid
  Rostroraja alba (Lacepède, 1803) -- valid -- bottlenosed skate, white skate
  Springeria folirostris Bigelow and Schroeder, 1951 -- invalid
  Sympterygia Müller and Henle, 1837 -- valid
  Sympterygia acuta Garman, 1877 -- valid
  Sympterygia bonapartii Müller and Henle, 1841 -- valid
  Sympterygia brevicaudata (Cope, 1877) -- valid
  Sympterygia lima (Poeppig, 1835) -- valid
   

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