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Boulenophrys  Fei, Ye and Jiang, 2016
Taxonomic Serial No.: 1094534

(Download Help) Boulenophrys TSN 1094534

 Taxonomy and Nomenclature
       
  Kingdom: Animalia  
  Taxonomic Rank: Genus  
  Synonym(s): Panophrys Rao and Yang, 1997
 
    Tianophrys Fei and Ye, 2016
 
  Common Name(s):    
       
  Taxonomic Status:    
  Current Standing: valid  
       
  Data Quality Indicators:    
  Record Credibility Rating: verified - standards met  
  Global Species Completeness: complete   
  Latest Record Review: 2025   
       

 Taxonomic Hierarchy
       
 KingdomAnimalia  – Animal, animaux, animals  
    SubkingdomBilateria  – triploblasts  
       InfrakingdomDeuterostomia   
          PhylumChordata  – cordés, cordado, chordates  
             SubphylumVertebrata  – vertebrado, vertébrés, vertebrates  
                InfraphylumGnathostomata   
                   SuperclassTetrapoda   
                      ClassAmphibia  – amphibiens, anfíbio, Amphibians  
                         OrderAnura  – Frogs, perereca, rã, sapo, crapauds, grenouilles, Toads  
                            FamilyMegophryidae Bonaparte, 1850  
                               SubfamilyMegophryinae Bonaparte, 1850  
                                  GenusBoulenophrys Fei, Ye and Jiang, 2016  
    Direct Children:  
                                     Species Boulenophrys acuta (Wang, Li and Jin in Li, Jin, Zhao, Lin, Wang and Pang, 2014)  
                                     Species Boulenophrys angka (Wu, Suwannapoom, Poyarkov, Chen, Pawangkhanant, Xu, Jin, Murphy and Che, 2019)  
                                     Species Boulenophrys anlongensis (Li, Lu, Liu and Wang, 2020)  
                                     Species Boulenophrys baishanzuensis (Wu, Li, Liu, Wang and Wu, 2020)  
                                     Species Boulenophrys baolongensis (Ye, Fei and Xie, 2007)  
                                     Species Boulenophrys binchuanensis (Ye and Fei, 1995)  
                                     Species Boulenophrys binlingensis (Jiang, Fei and Ye in Fei, Hu, Ye and Huang, 2009)  
                                     Species Boulenophrys boettgeri (Boulenger, 1899)  
                                     Species Boulenophrys brachykolos (Inger and Romer, 1961)  
                                     Species Boulenophrys caobangensis (Nguyen, Pham, Nguyen, Luong and Ziegler, 2020)  
                                     Species Boulenophrys caudoprocta (Shen, 1994)  
                                     Species Boulenophrys changyangensis Li, Shi, Liu, Zhao, Gao and Wang, 2025  
                                     Species Boulenophrys cheni (Wang and Liu in Wang, Zhao, Yang, Zhou, Chen and Liu, 2014)  
                                     Species Boulenophrys chishuiensis (Xu, Li, Liu, Wei and Wang, 2020)  
                                     Species Boulenophrys congjiangensis (Luo, Wang, Wang, Lu, Wang, Deng and Zhou, 2021)  
                                     Species Boulenophrys daiyunensis (Lyu, Wang and Wang in Lyu et al., 2021)  
                                     Species Boulenophrys daoji (Lyu, Zeng, Wang and Wang in Lyu et al., 2021)  
                                     Species Boulenophrys daweimontis (Rao and Yang, 1997)  
                                     Species Boulenophrys daxuemontis Liu, Li, Cheng, Wei, Wang and Cheng, 2025  
                                     Species Boulenophrys dongguanensis (J. Wang and Y. Y. Wang in J. Wang et al., 2019)  
                                     Species Boulenophrys dupanglingensis Xiao and Mo, 2025  
                                     Species Boulenophrys elongata Zeng, Wang, Chen, Xiao, Zhan, Li and Lin, 2024  
                                     Species Boulenophrys fanjingmontis (Zhang, Liang, Ran and Shen, 2012)  
                                     Species Boulenophrys fansipanensis (Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong and Rowley, 2018)  
                                     Species Boulenophrys fengshunensis Wang, Zeng, Lyu and Wang, 2022  
                                     Species Boulenophrys frigida (Tapley, Cutajar, Nguyen, Portway, Mahony, Nguyen, Harding, Luong and Rowley, 2021)  
                                     Species Boulenophrys gaolanensis Song, Wang, Qi, Wang and Wang, 2024  
                                     Species Boulenophrys hengshanensis Qian, Hu, Mo, Gao, Zhang and Yang, 2023  
                                     Species Boulenophrys hoanglienensis (Tapley, Cutajar, Mahony, Nguyen, Dau, Luong, Le, Nguyen, Nguyen, Portway, Luong and Rowley, 2018)  
                                     Species Boulenophrys huangniushiensis Wang, Wu, Song, Huang, Zhuo and Wang, 2025 – Huangniushi Horned Toad 
                                     Species Boulenophrys hungtai Wang, Zeng, Lyu, Xiao and Wang, 2022  
                                     Species Boulenophrys insularis (Wang, Liu, Lyu, Zeng and Wang, 2017)  
                                     Species Boulenophrys jiangi (Liu, Li, Wei, Xu, Cheng, Wang and Wu, 2020)  
                                     Species Boulenophrys jingdongensis (Fei and Ye in Fei, Ye and Huang, 1983)  
                                     Species Boulenophrys jinggangensis (Wang in Wang, Zhang, Zhao, Sung, Yang, Pang and Zhang, 2012)  
                                     Species Boulenophrys jiulianensis (J. Wang, Zeng, Lyu and Y. Y. Wang in Wang et al., 2019)  
                                     Species Boulenophrys kuatunensis (Pope, 1929)  
                                     Species Boulenophrys leishanensis (Li, Xu, Liu, Jiang, Wei and Wang, 2019)  
                                     Species Boulenophrys liboensis (Zhang, Li, Xiao, Li, Pan, Wang, Zhang and Zhou, 2017)  
                                     Species Boulenophrys lichun Lin, Chen, Li, Peng, Zeng and Wang, 2024  
                                     Species Boulenophrys lini (Wang and Yang in Wang, Zhao, Yang, Zhou, Chen and Liu, 2014)  
                                     Species Boulenophrys lishuiensis (Wang, Liu and Jiang in Wang, Liu, Jiang, Jin, Xu and Wu, 2017)  
                                     Species Boulenophrys lushuiensis (Shi, Li, Zhu, Jiang, Jiang and Wang, 2021)  
                                     Species Boulenophrys minor (Stejneger, 1926)  
                                     Species Boulenophrys mirabilis (Lyu, Wang and Zhao in Lyu, Li, Zeng et al., 2020)  
                                     Species Boulenophrys mufumontana (J. Wang, Lyu and Y. Y. Wang in J. Wang et al., 2019)  
                                     Species Boulenophrys nankunensis (J. Wang, Zeng and Y. Y. Wang in J. Wang et al., 2019)  
                                     Species Boulenophrys nanlingensis (Lyu, J. Wang, Liu and Y. Y. Wang in J. Wang et al., 2019)  
                                     Species Boulenophrys obesa (Wang, Li and Zhao in Li, Jin, Zhao, Liu, Wang and Pang, 2014)  
                                     Species Boulenophrys ombrophila (Messenger and Dahn in Messenger et al., 2019)  
                                     Species Boulenophrys omeimontis (Liu, 1950)  
                                     Species Boulenophrys palpebralespinosa (Bourret, 1937)  
                                     Species Boulenophrys pepe Wang and Zeng, 2024  
                                     Species Boulenophrys puningensis Wang, Zeng, Lyu, Xiao and Wang, 2022  
                                     Species Boulenophrys qianbeiensis (Su, Shi, Wu, Li, Yao, Wang and Li, 2020)  
                                     Species Boulenophrys rubrimera (Tapley, Cutajar, Mahony, C. T. Nguyen, Dau, T. T. Nguyen, Luong and Rowley, 2017)  
                                     Species Boulenophrys sangzhiensis (Jiang, Ye and Fei, 2008)  
                                     Species Boulenophrys sanmingensis (Lyu and Wang in Lyu, Zeng, Wang, Liu, Huang, Li and Wang, 2021)  
                                     Species Boulenophrys shimentaina (Lyu, Liu and Wang in Lyu, Li, Zeng, Zhao, Liu, Guo and Wang, 2020)  
                                     Species Boulenophrys shuichengensis (Tian and Sun, 1995)  
                                     Species Boulenophrys shunhuangensis (Wang, Deng, Liu, Wu and Liu, 2019)  
                                     Species Boulenophrys spinata (Liu and Hu in Hu, Zhao and Liu, 1973)  
                                     Species Boulenophrys tongboensis (Wang and Lyu in Lyu, Zeng, Wang, Liu, Huang, Li and Wang, 2021)  
                                     Species Boulenophrys tuberogranulata (Shen, Mo and Li in Mo, Shen, Li and Wu, 2010)  
                                     Species Boulenophrys wugongensis (J. Wang, Lyu and Y. Y. Wang in J. Wang et al., 2019)  
                                     Species Boulenophrys wuliangshanensis (Ye and Fei, 1995)  
                                     Species Boulenophrys wushanensis (Ye and Fei, 1995)  
                                     Species Boulenophrys xiangnanensis (Lyu, Zeng and Wang in Lyu, Li, Zheng, Zhao, Liu, Guo and Wang, 2020)  
                                     Species Boulenophrys xianjuensis (Wang, Wu, Peng, Shi, Lu and Wu, 2020)  
                                     Species Boulenophrys xuefengmontis Lyu and Wang, 2023  
                                     Species Boulenophrys yangmingensis (Lyu, Zeng and Wang in Lyu et al., 2020)  
                                     Species Boulenophrys yaoshanensis Qi, Mo, Lyu, Wang and Wang, 2021  
                                     Species Boulenophrys yezhongensis Liu, Feng, Shen, Li, Cheng, Wei, Wang and Su, 2025  
                                     Species Boulenophrys yingdeensis Qi, Lyu, Wang and Wang, 2021  
                                     Species Boulenophrys yunkaiensis Qi, Wang, Lyu and Wang, 2021  
       

 References
       
  Expert(s):    
  Expert:    
  Notes:    
  Reference for:    
       
  Other Source(s):    
  Source: Amphibian Species of the World 6.2 2025-09-15, database (version 6.2)  
  Acquired: 2025   
  Notes: Frost, Darrel R. 2025. Amphibian Species of the World: an Online Reference. Version 6.2 (15 September, 2025). lectronic Database accessible at https://amphibiansoftheworld.amnh.org/index.php American Museum of Natural History, New York, USA   
  Reference for: Boulenophrys   
       
  Source: Amphibian Species of the World: an Online Reference v6, database (version 6.0)  
  Acquired: 2019   
  Notes: Frost, Darrel R. 2019. Amphibian Species of the World: an Online Reference. Version 6 (12 September, 2019). Electronic Database accessible at http://research.amnh.org/herpetology/amphibia/index.html American Museum of Natural History, New York, USA   
  Reference for: Boulenophrys   
       
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